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Sedge swollen (Carex rhynchophysa) 3

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Description: The fruit (often incorrectly called a nut) is dry, single-seeded, not opening, formed by the upper ovary of a three-membered cenocarpous gynoecium and completely enclosed in a closed sac. The sac in shape is almost round or round-ovate, 5-6.5 mm long, obscurely trihedral, swollen, almost round in cross section, with few long, not protruding thin veins, thin-leathery, shiny, smooth, reddish-straw yellow, rounded wedge-shaped narrowed at the base, sharply turning into a short leg, sharply narrowed at the top to an elongated, equal to about 1/3 of the length of the sac, about 1.5-2 mm long, narrow cylindrical, at the top a shallow wedge-shaped dissected biconvex with generally good p zvitymi prongs to 0.5 mm. The fruit is narrowly obovate, trihedral, with a nose curved at the base, brown.

Label: Test tube collection: 1628

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Genus Puhonos - Trichophorum

Trichophorum caespitosum (L.) Hartm. - Downy soddy. Everywhere in the zone of boreal elfin and southern hypoarctic tundra. Not rarely. The shores of lakes, potholes on terraces, sedge marshes, sphagnum sedge tundra. Circumpolar alpine-hypoarctic bog species. From the northern part of the Koryak Upland is not known, it is indicated only for the hall. Korfa and the river basin Oklan, as well as for the river basin. Anadyr.

Upper river Khatyrka: intermountain trench near Mount Evlanyr, along the lake, 07/20/1993, Middle and lower reaches of the river. Great: 15 km above the mouth of the river. Tamvatvaam, a swamp at the head of the stream, 2.08.1993, Upper river. Chiranay: p. Mayolgykonvaam in the middle course, duct in a high floodplain, 07/06/1995, The middle course of the river. Chiranay: the vicinity of Lake Good, tundra on the terrace, Mozhazhina, 07/31/1995, Upper river. Nygchekveiem: against Mount George (turn), the edge of the sedge hummocky and shrub tundra on the terrace, 07.24.1996, the Middle course of the river. Nygchekveem: in the range of the mountain Nygchek - lake. The eternal sedge swamp by the moraine lake, 08/03/1996.

Trichophorum caespitosum - Downy sod

Eleocharis acicularis - Needle swamp

Eleocharis palustris - Swamp marsh

Rod Mireland - Eleocharis

Eleocharis acicularis (L.) Roem. et Schult. - Needle swamp. Everywhere in the zone of boreal elfin and southern hypoarctic tundra. Not rarely. Floodplain sands, especially silty, along the channels on terraces, bottoms of drained moraine lakes. Circumpolar boreal marsh species.For the northern part of the Koryak Upland, it was not given before.

Middle and lower river. Great: the mouth of the river. Tamvatvaam, floodplain sands, 08/03/1993, the mouth of the river. Etchinku, a quiet backwater, sedge meadow along a silty sandy beach, 08/07/1993, oil refinery base area, opposite Peschany island, 4th terrace, silty duct, 08/17/1995, in the same place, duct on the terrace, 08/10/1993, Average course of the river . Nygchekveem: moraine near the mountain Nygchek, Lake. Eagle, on the bottom of the drained lake, 08/05/1996, lake. Eternal, rocky-silty sandbank. 07/29/1996.

Eleocharis palustris (L.) Roem. et Schult. = E. intersita Zinserl. - Swamp swamp. Everywhere in the zone of boreal elfin and southern hypoarctic tundra. Infrequently. Eurasian-North American hypoarctoboreal bog species. In all likelihood, a race lives in Northeast Eurasia, which can be considered as an independent subspecies.

The average course of the river. Chiranay: the area of ​​the mouth of the river. Shallow, silty duct, 07.29.1995.

Eriophorum brachyantherum - Short-anther bush

Rod Fluffy - Eripohorum

Eriophorum brachyantherum Trautv. et C.A. Mey. - Short-boot carapace. Seldom. Drawdowns and depressions on fluvioglacial deposits. Circumpolar hypoarctic swamp-tundra species. In all likelihood, this is an ecological variation. E. vaginatum L. It differs then by its very loose turfs - half-chunks. The sign of bending-inbending scales is a bad sign. There are no collections of this species from the Koryak Upland (Arctic flora of the USSR).

Upper river Khatyrka: mouth of the creek. Maisky, high terrace, thermokarst depressions, 07/12/1993.

Eriophorum medium Anderss. - Fluffy medium. Everywhere. Sporadically. It is rarely abundant. Swamps and moist tundra. Circumpolar hypoarctic bog species. Very slightly different from E. russeolum. There are very few collections of this species from the Koryak Upland.

Upper river Khatyrka: ruch. Maysky, floodplain, 07/12/1993, ibid., High terrace, sphagnum sedge swamp with rosemary, 07/16/1993, thermokarst depressions on a high terrace, 07/12/1993, Upper river. Chiranay: mouth of the river. Mayolgykonvaam, high floodplain, high bushes, 07/07/1995, in the same place, floodplain shrubs along the channels, 07/06/1995.

Eriophorum polystachyon L. aggr. ssp. subarcticum V. vassil. - Multi-spike gun

Eriophorum polystachyon L. ssp. komarovii (V. Vassil.) Galanin comb. prov. - Fluffy multi-spike Komarova. Everywhere. Pretty common. Kochkarnye tundra, swamps, potholes, sometimes in pike meadows. North American-East Asian boreal bog species. The plant is more subtle compared to a typical subspecies, the leaves are curled.

Tamvatney mountains: Martian mountain, gravelly-loamy plume, sedge tundra, 07.28.1994, Tumannaya saddle, tundra with elfin, 07.27.1994, Upper river. Chiranay: mouth of the river. Mayolgykonvaam, high floodplain, high shrubs, 07/07/1995, The middle course of the river. Chiranay: p. The winding, cedar-lined hummock tundra on the hill, the spur, 2.07.1995, at the trigger point 78 m, tuberous shrubby tundra, 07.28.1995, Upper course of the river. Nygchekveem: against Mount George (turn), pike meadow at the mouth of the stream. Patrol, 07.26.1996, Surroundings of the city of Anadyr: 15th km, cliff, 06.29.1993.

Eriophorum polystachyon L. aggr. ssp. subarcticum V. vassil. - Multi-spike gun. Everywhere. Pretty common. Marshes, hummocky tundra, sedge wet meadows, along the banks of moraine lakes. Circumpolar arctoboreal tundra-bog species of wide amplitude. Powerful plant, wide leaves. There are very few collections of this species from the Koryak Upland.

River basin Tamvatvaam: the watershed of the Tumgytveem and Razliv rivers, sedge-mixed grass eutrophic swamp, 07/19/1994, the source of the river. Spill, shore of a moraine lake, 07/19/1994, The middle course of the river. Chiranay: p. Sinuous, spalls in the hummock tundra with cedar dwarf, 2.07.1995, surroundings of lake. Good, high terrace, kochkarnaya tundra with dwarf, 07/30/1995, Upper river. Nygchekveem: against Mount Janranai, sedge meadow on the shore of Lake Nizinnoe, 07/19/1996, The middle course of the river. Nygchekveem: in the range of the mountain Nygchek - lake. Eternal, thermokarst failure on the moraine, sedge swamp, 07/31/1996, lake with cyclut thickets, 08/03/1996.

Eriophorum russeolum Fries. - Fluffy reddish. Everywhere. Usually. Swamps, pike meadows, moist tundra. Circumpolar hypoarctic bog species. Bracts of scales black with a wide white border around the edge. For the northern part of the Koryak Upland is given for the first time.

Upper river Khatyrka: ruch.Maisky, high terrace, oligotrophic swamp along a stream, 07/16/1993, River basin. Tamvatvaam: the lower reaches of the river. Tundra, icy valley, 07/23/1994, watershed of the Tumgytveem and Razliv rivers, sedge-mixed grass eutrophic swamp, 07/19/1994, shore of a moraine lake at the head of the river. Spill, 07/19/1994, Upper course of the river. Chyrynai: Chyrynai mountains, swamp on a train, pike meadow, 07/22/1995, the upper river. Mayolgykonvaam, the icy section of the valley, 07/17/1995, Mount Ugryumaya, swamp in the valley of the river. Vilyui, 07.15.1995, Sources of the river. Nygchekveem, Koryak ridge: the mouth of the stream. find, swamp on the fluvioglacial terrace, 07/11/1996, moss-mixed grassland on the mountain plume, 07/06/1996, Upper course of the river. Nygchekveiem: against Mount George (turn), sedge swamp on the moraine, 07.24.1996, the Middle course of the river. Nygchekveem: in the range of the mountain Nygchek and Lake. Eternal, swamp on the floodplain terrace, 07/29/1996, in the same place, sedge swamp on the floodplain terrace, 07/29/1996, Outskirts of the city of Anadyr: Mount Dionysia, plume, raw tundra, 2.07.1993.

Eriophorum scheuchzeri Hoppe - Sheikhtser Fluff. Everywhere. Quite often, sometimes a massive background look. Swamps, pike meadows, moist tundra. Circumpolar arctoalpine tundra-bog species. For the Koryak upland in the Arctic flora of the USSR, fees of this species are not given.

Upper river Khatyrka: ruch. Maisky, floodplain, 07/12/1993, Tamvatney mountains: sand spit p. Tamvatvaam, silted pike meadow, 07/27/1994, shore of a flowing lake in the floodplain of the Tamvatvaam river, 07/29/1994, River basin. Tamvatvaam: the middle course of the river. Tundra, icy section of the valley, 21.07.1994, Upper course of the river. Chiranay: ruch. Dry, eutrophic swamp in the valley, spar between hillocks, massively, 07/11/1995, near the Chyrynai mountains, a plume of a hill, among erniks, 07/23/1995, in the same place, a wet place on a plume of a dump among yerniks and elfin, 07/23/1995, Middle course R. Chiranay: the vicinity of Lake Good, kochkarnaya tundra on the terrace, cross-country road, 07/30/1995, the mouth of the river. Openwork, 1st floodplain terrace, 6.08.1995, Upper course of the river. Nygchekveem: near the mouth of the river. Kelmyveem, silted duct in the floodplain of the river, 07/16/1996, against Mount George (turn), the cone of removal of the stream into the floodplain of the river, background view, 07/27/1996, against Mount Janranay, sedge meadow along the shore of the lake. Lowland, on pebble, 07/19/1996, Surroundings of the city of Anadyr: Mount Michael, slope, along the stream, 07/04/1993.

Eriophorum vaginatum L. - Vaginal fluff. Only in the zone of boreal dwarfs and southern hypoarctic tundras. Infrequently. Swamps, pike meadows, hummock tundra. Circumpolar hypoarctic tundra species. There were no collections of this species from the northern part of the Koryak Upland.

Upper river Khatyrka: ruch. Maisky, thermokarst depressions on a high terrace, 07/12/1993, River basin. Tamvatvaam: the middle course of the river. Tumgytvey, shrubs of the middle floodplain, 07/16/1994, watershed of the Tumgytvei and Razliv rivers, sedge swamp, 07/19/1994, Middle course of the river. Chiranay: the vicinity of Lake Good, high terrace, kochkarnaya tundra with elfin, 07/30/1995, p. Winding, top of a bump, elfin tundra, 2.07.1995.

Eriophorum polystachyon - Multi-spikelet

Eriophorum vaginatum - Vaginal fluff

Rod Sedge - Carex

Сarex aquatilis Wahlenb. - Sedge water. Everywhere. Usually. Swamps, pits, wet and hummock tundra, lake shores. Circumpolar boreal marsh species. For the northern part of the Koryak Upland is given for the first time.

Middle and lower river. Velikaya: surroundings of the oil refiner depot, the spur in the low tundra, 08.1993, 2 km below the translucent sign 28 m, surroundings of Lake Vase, hummock tundra on a high terrace, in the spar, 08.15.1995, ravine on a high terrace, 08.3.1996, River basin. Tamvatvaam: the watershed of the Tumgytveem and Razliv rivers, sedge-mixed grass swamp, 07/19/1994, p. Tamvatvaam 10 km above the mouth of the river. Nauchirynai, the bottom of a dried lake in the tundra, 2.08.1994, Upper course of the river. Chiranay: Mount Gloomy, swamp in the valley of the river. Vilyui, 07.15.1995, valley of the brook. Dry, eutrophic tuberous bog with pits, 07/11/1995, Middle course of the river. Chiranay: the vicinity of Lake A good ridge-ridge structure in the kochkarnaya tundra, 1.08.1995, in the same place, a swamp along the edge of the lake, 07.31.1995, Winding, hillside, bottom of the drain trough, 2.07.1995, Upper river. Nygchekveiem: against Mount George (turn), sedge swamp with peat mounds on the terrace, 07.24.1996, The middle course of the river. Nygchekveem: the vicinity of Lake. The eternal, floodplain terrace of the river, the edge of the sedge swamp, 07/29/1996, Mount Nychek, lake. Lonely Owl, in the water and offshore, 08/05/1996, Surroundings of the lake.Mainz: the western shore of the lake against Tumanny Island, a key swamp on the moraine terrace, 07/20/1996.

Carex alticola Poplavsk. ex Sukacz. - Sedge Alpine. East Siberian alpine-tundra species. News for the Koryak Upland as a whole, the closest locations in Okhotia south of the river. Pits. Only one specimen was collected, randomly along with other species.

The average course of the river. Nygchekveem: the vicinity of Lake. Smooth, shrubby tundra on the floodplain terrace, 06/28/1996.

Carex amgunensis Fr. Schmidt. - Sedge Amgun. Seldom. Eurasian boreal forest species. Deviates to C. globularis L., from which it differs by more compact inflorescences and in that it forms rather dense sods. News for the Koryak Highlands and Chukotka.

The average course of the river. Nygchekveem: in the alignment Lake Eternal - Mount Nygchek, ernikovaya tundra on the floodplain terrace of the river, 08/03/1996, the beach of Lake Eternal, 07/29/1996, the surroundings of the former village. Maino-Gytkino, conflagration on the fluvioglacial terrace, 08/09/1996.

Carex anthoxanthea Presl. - Sedge spikelet. Swamps on mountain plumes and moraine depressions. Beringian arctic tundra view. Known from the vicinity of the village. Providence and from the Commander Islands. For Koryakskoye highlands not previously given.

Upper river Khatyrka: ruch. Maisky, thermokarst depressions on the high moraine terrace of the river, 07/12/1993, floodplain of the river, 07/12/1993, Sources of the river. Nygchekveem, Koryak Range: creek. Find, mixed-moss swamp on a train, July 6, 1996.

Carex appendiculata (Trautv. Et Mey.) Kuk. - sedge attached. Everywhere. Very ordinary, sometimes abundant and dominant. Raw meadows and swamps. East Siberian-Far Eastern boreal species. For the Northern Koryak Highlands, it was not given before. The upper pistillate spikelet is often with stamen flowers. Typical specimens have yellow pubescent roots.

Upper river Khatyrka: mouth of the creek. Maisky, thermokarst depressions on a high moraine terrace, 07/12/1993, ancient river terrace, along a brook, 07/16/1993, along a brook in a narrow ravine on a high river terrace, 07/16/1993, a swamp along a stream on a high terrace, 07/16/1993, Pool R. Koyverelyan: p. Vilyumeikuul, intermountain hollow, first floodplain terrace, 07.24.1993, ibid., Willow on the floodplain terrace, 07.24.1993, mouth of the river. Koyverelyan, silty duct in the floodplain, alder, 07/29/1993, Middle and lower reaches of the river. Great: sandy island below the mouth of the river. Chiranay, willow from S. udensis along the channel, 08/12/1995, River basin. Tamvatvaam: the lower reaches of the river. Tundra, icy valley, 07.23.1994, the middle course of the river. Tumgytveem, glade in the dwarf thickets on the floodplain terrace, 07.17.1994, the mouth of the river. Tundra, old sparse poplar forest, duct, 07/24/1994, Upper course of the river. Chiranay: surroundings of Mount Ugryumaya, floodplain terrace of the river. Vilyui, 07.15.1995, valley of the river. Vilyui, swamp, 07.15.1995, the mouth of the river. Mayolgykonvaam, high moraine terrace, 07/04/1995, The middle course of the river. Chiranay: raw duct in the floodplain alder, forms large hummocks, 08/05/1995, the vicinity of Lake. Good, willow from S. pulchra on the terrace, 07/31/1995, in the same place, the shore of the drying up logs in the tundra, 07/31/1995, raw duct in the floodplain, hummock in alder, 08/08/1995, Upper course of the river. Nygchekveem: against Mount George (turn), mixed willows from S. pulchra on the terrace, 07.24.1996, against Mount Janranai, shore of the lake. Low sedge meadow, 07/19/1996, The middle course of the river. Nygchekveem: the vicinity of Lake. Eternal, drained lake near the river. Silver, 07/29/1996, against the lake. Gemini, floodplain terrace near the river. Silver, 08/03/1996.

Carex appendiculata (Trautv. Et Mey.) Kuk. x C. saxatilis L. A very strange plant, powerful, well-leaved, combining the characteristics of both species, but more like C. appendiculata. Spikelets, especially female ones, are clearly underdeveloped. This is even more noticeable with the good development of leaves and stems.

Middle and lower river. Great: 6 km above the weather station, meadow along the canal in the floodplain, 08/15/1995.

Carex atrofusca - sedge black and brown

Carex atrofusca Schkuhr. - sedge black and brown. Sporadically not rare. Tundra, cedar dwarfs and mixed lawns in the mountains of the main composition. Circumpolar arctoalpine mountain-bog species. In some individuals, the sacs and bracts are not black, but brown. For the Koryak Upland as a whole, there is one gathering from the south of Kamchatka and from the lower Penzhina.

Tamvatney mountains: the lower reaches of the stream.Camp, lawn for grass, 07/26/1994, the lower part of the Martian mountain slope, spotted herb tundra with elfin, 07/27/1994, Tumannaya mountain, tundra with elfin on the saddle, 07/27/1994,

Carex bicolor Bell. ex Allioni - Two-color sedge. Seldom. Pebble gravels, silty ducts, loamy spots in the hummock tundra with thickets of cedar dwarf elf. Arctoalpine species with extremely intermittent distribution. The nearest location - on the river. Palyavaam and in the east of the Chukchi Peninsula. For the Koryak Highlands is given for the first time, the closest location on the river. Palyavaam and in the east of the Chukchi Peninsula, where it is found on carbonate pebbles. Small sedge, plants 6-8 (10) cm high, rhizome, spikelets small, crowded, upper with a female, but with male colors, stigma 2. White sacs from powdery powdery coating, without spouts.

The average course of the river. Chiranay: p. Winding, moraine ridge, loamy spots in the hummock tundra with thickets of cedar dwarf, 2.07.1995, mouth of the river. Openwork, gravel of a floodplain, 5.08.1995 (strongly avoids C. eleusinoides Turcz. ex Kunth, but differs from the latter by the absence of a nose in the sacs and a set aside lower spikelet), Upper course of the river. Nygchekveem: the vicinity of Lake. Smooth, muddy duct in the floodplain of the river, 1.07.1996,

Carex bigelowii ssp. ensifolia (Gorodk.) Holub. = C. ensifolia Turcz. ex Krecz. - Sedge mesaceous. Everywhere. Usually. Tundra, swamps, sedge meadows. Sometimes very plentiful. Siberian-Mongolian arctoalpine mountain tundra species. Roots with yellow pubescence.

Upper river Khatyrka: ruch. Maisky, thermokarst depressions on a high terrace, 07/12/1993 (four leaves), intermountain hollow near Mount Evlanir, high floodplain terrace, 07/20/1993, River basin. Tamvatvaam: the middle course of the river. Tumgytvey, dividing log, sedge bushes, 07/17/1994, shrubs of the middle floodplain along the river. Tumgytvey, 07.16.1994, Upper course of the river. Chyrynai: a plume of the slope of the Chyrynai mountains, raw springs, 07/22/1995, Sources of the river. Nygchekveem, Koryak Range: creek. Finding, wet swampy meadow on a plume, usual, 07/06/1996.

Carex bigelowii ssp. rig>

The surroundings of Lake Mainz: the western shore of the lake against the island of Tumanny, a small-pebble beach, near the water in large numbers, 07/20/1996.

Carex bonanzensis Britt. - Sedge Bonancine. Seldom. Willows on streams, tuberous swamps with springs, icy river valleys. Alaskan-Asian boreal-hypoarctic species of riverine ecotopes. For the Koryak Highlands, in general, news, but there are fees for Anadyr and Penzhin.

River basin Tamvatvaam: the lower reaches of the river. Tundra, icy valley, 07.23.1994, Upper river. Chiranay: ruch. Dry, tuberous swamp with pits in the valley, 07/11/1995, Upper river. Nygchekveem: against Mount George (turn), willows from S. alaxensis forbs along the brook on the terrace, 07.24.1996.

Carex capillaris L. = C. chlorostachys Steven - Hairy sedge. Everywhere. Not rarely. Meadows, willows, tundra, often along shallow watercourses. Circumpolar (?) Hypoarctoalpine mountain tundra species. News for the Koryak Highlands. In our plants, the noses of the pouches are smooth, very rarely jagged. The plants are rather tall, up to 30 cm high.

Upper river Khatyrka: thermokarst depressions on a high glacier terrace, 07/12/1993, Tamvatney mountains: curtain sedge tundra and mixed lawn along a mountain stream, 07/27/1994, saddle of the Tumannaya Mountain, 07/27/1994, River basin. Tamvatvaam: p. Tundra at the mouth, floodplain terrace, glade among thickets of elfin trees, 07.24.1994, in the same place, plume, herbaceous and shrub tundra, 07.24.1994, Upper course of the river. Chyrynai: saddle on Mount Gloomy, loamy spots, 07/13/1995, p. Mayolgykonvaam in the middle course, duct in a high floodplain, mixed willow, July 6, 1995, the mouth of the river. Elgiveem, Mount Chelka, sedge tundra along the plume, 07/19/1995, surroundings of Ugryumaya mountain, floodplain terrace of the Vilyuy river, bushes, 07/15/1995, mixed shrubs on the hillside, 07/23/1995, The middle course of the river. Chyrynay: at the trigger point 78 m, shrubby tuberous tundra on a high terrace, 07.28.1995, Upper course of the river. Nygchekveem: mouth of the river. Ichguigeynei, ledge of a high terrace, nival hollow on the terrace, 06/30/1996, against Mount George (turn), willow forbs along the brook on the terrace, 07/24/1996, The middle course of the river. Nygchekveem: against the lake. Gemini, p. Silver, ernik tundra on the terrace, 08/03/1996.

Сarex capitata L. - Capitate sedge. Everywhere in the zone of boreal dwarfs. Not rarely. On shallow watercourses.Circumpolar boreal marsh species. For the northern part of the Koryak Upland is given for the first time.

Upper river Khatyrka: high river terrace, dry lichen tundra with willows, 07/16/1993, Tamvatney mountains: low floodplain terrace, bank of the canal, 08/29/1994, River basin. Tamvatvaam: Tundra river at the mouth, drain trough in the tundra, willows from S. pulchra, 07.24.1994, ibid., Description 32, 07.24.1994, Upper course of the river. Chiranay: Ugryumaya mountain, the valley of the Vilyuy river, Ivnyaki, 07/15/1995, the mouth of the Elgeveem river, a drain hollow on a gentle slope, 07/19/1995, Upper course of the Nygchevei river: the vicinity of Lake Smooth, silty ducts in the floodplain of the river, 1.07.1996, against Mount George, turn, stream on the terrace, willows from S. alaxensis, 24.07.1996.

Carex caespitosa L. - sedge

Carex cespitosa L. - sedge. Everywhere in the zone of boreal dwarfs. Not rarely. Meadows, shrubs, tundra. Palearctic boreal marsh species. For the Koryak Upland is given for the first time, it is found along Anadyr and Omolon, one collection from the southern Koryak Upland is known. It seems to us that C. appendiculata, C. schmidtii, C. wiluica, C. minuta should be considered as subspecies and even varieties C. caespitosa L. In our area, all these subspecies form an inseparable morphological continuum, and specific individuals can most often be determined only up to C. caespitosa s.l. Most features C. caespitosa s.str. characteristic of plants collected in the river basin. Chiranay. Perhaps this is somehow connected with the local meadows and hyperbasites.

Tamvatney mountains: low floodplain terrace of the river. Tamvatvaam, shrub tundra, 07/29/1994, Upper river. Chyrynai: against the Chyrynai mountains, a swampy meadow along a stream, 07/21/1995, the vicinity of Mount Ugryumaya, the first floodplain terrace of the river. Vilyui, bushes, 07.15.1995, the mouth of the river. Maiolgykonvaam, a grass meadow in a hollow on a high fluvioglacial terrace, 07/11/1995, Middle course of the river. Chyrynay: surroundings of the trigger point 78 m, dry groove on the terrace, meadows in willows, 07.28.1995, the mouth of the river. Openwork, raw duct in the floodplain alder, 08/05/1995.

Carex cinerea Poll. = C. canescens auct - Gray sedge. Everywhere. Not rarely. Swamps, hollows of runoff. Circumpolar arctoboreal bog species. Previously given for the northern part of the Koryak Upland only for Ugolnaya Bay. There are fees from the southern part of the Koryak Highlands.

Upper river Khatyrka: mouth of the creek. Maisky, thermokarst depressions on a high moraine terrace, 07/12/1993, swamp along a stream with peat mounds, high terrace, 07/16/1993, River basin. Tamvatvaam: p. Tundra, drain hollow on the slope of the boulder, willows from S. pulchra, 04/27/1994, watershed of the Tumgytveem and Razliv rivers, sedge-mixed grass swamp, 07/19/1994, Upper course of the river. Chiranay: valley brook. Dry, eutrophic tuberous bog with pits, 07/11/1995, surroundings of Mount Ugryumaya, swamp in the valley of the river. Vilyui, 07.15.1995, Surroundings of Lake Mainz: west coast against Tumanny Island, a key swamp on a moraine terrace, 07/20/1996.

Carex cryptocarpa C.A. Mey. - sedge crypto-fertile. Everywhere. Silted banks of water bodies, along streams, floodplain willows, meadows, gravels, sedge marshes. North American-East Asian Arctic boreal meadow-bog species. For the northern part of the Koryak Upland is given for the first time, although there are many fees from neighboring places. Our plants have bags with more or less clearly visible veins. The upper two pistil spikelets at the end are always with male flowers. Sometimes all female spikelets with male flowers. The base of the shoots is dark cherry, leaves up to 0.5 cm wide or more.

Upper river Khatyrka: intermountain basin near Mount Evlanyr, dominant by the stream, 07/20/1993, River basin. Koyverelyan: near the mouth of the river. Koyverelyan, willows of the first floodplain terrace, 07.24.1993, Middle and lower reaches of the river. Great: 6 km above the weather station, meadow on a silty sandy beach, 08/15/1995, in the same place, meadow along a temporary channel, 08/15/1995, Tamvatney mountains: sand spit on the river. Tamvatvaam, silted area, pike meadow, 07.27.1994, Middle course of the river. Chiranay: against the river. Shallow at the mouth, under the ledge of a high terrace, silted duct, 07/29/1995, near the mouth of the river. Openwork, sandy shore, 08/06/1995, silty duct on the banks of the river. Openwork, 08/06/1995, The sources of the river. Nygchekveem, Koryak ridge: the mouth of the stream. Nakhodka, swamp with a stream on a high floodplain terrace of the river, 07/11/1996, Upper course of the river. Nygchekveem: surroundings of Janranai Mountain, the shore of Lake Lowland, plentiful, 07/19/1996, near the mouth of the river. Kelmyveem, swamp in the river valley, 07/16/1996, the vicinity of the lake.Smooth, floodplain pebbles, 07/03/1996, Middle course of the river. Nygchekveem: the vicinity of Lake. Eternal, p. Silvery, silted beach, 07.29.1996, high floodplain p. Nygchekveem, the shore of the channels against the lake. Eternal, 07/31/1996, floodplain terrace, sedge swamp, 07/29/1996, Surroundings of the lake. Mainz: Lake Gytgykay, the bank of the river at its source from the lake, Outskirts of Anadyr: Mount Dionysia, Second River, along the watercourse, 07.1993.

Carex delicata C. B. Clarke = C. karoi (Freyn) Freyn - Sedge tender. Seldom. Loamy spots in the tundra, lakeside. American-Asian shrub-bog species. In all likelihood, our plants belong to a race described as C. boecheriana A. et D. Love et Raymond, which is endemic to Greenland.

Middle and lower reaches of the Chyrynai River: the vicinity of Lake Good, willow from Salix pulchra along a stream flowing from the lake, a loamy spot in the eutrophic tundra, 1.08.1995, Upper river. Nygchekveem: against the mountain Yanranay, the shore of the lake. Lowland, 07/19/1996.

Carex dichroa (Freyn) V. Krecz. - Sedge bicolored. Everywhere in the zone of boreal dwarfs. Not rarely. Meadows, willows, shores of watercourses. Asian boreal meadow-bog species. Forms dense sods (almost bumps). Sacks of typical plants are light, yellow-green, scales are light brown, female spikelets are up to 3 cm long, hanging on long peduncles, the stems are rough. Our plants shy away from C. saxatilis L., although in general, undoubtedly belong to this species. For Chukotka and the Koryak Uplands is given for the first time, the closest locations are in the Kolyma Uplands. Light and dark colored forms are distinguished.

Tamvatney mountains: low floodplain terrace of the river. Tamvatvaam, a shrub tundra, forms powerful bumps, 07/29/1994, River basin. Tamvatvaam: p. Tundra at the mouth, 07.24.1994, Upper course of the river. Chiranay: mouth of the river. Mayolgykonvaam, high floodplain, willows, 07/07/1995, The middle course of the river. Chiranay: the vicinity of Lake Good, ledge of a high terrace, meadow, 1.08.1995, near the mouth of the river. Openwork, sandy river bank. Canyon, 08/08/1995, The upper course of the river. Nygchekveem: against the mountain of George (turn), shrubs on the terrace, 07.24.1996, the vicinity of Lake. Smooth, sedge meadow along the lake, 06/29/1996, The middle course of the river. Nygchekveem: the target is Nygchek Mountain - Lake Eternal, floodplain terrace on the right bank of the river, bank of the groove, 07/31/1996, in the same place, bushes of the first floodplain terrace, 07/29/1996, in the same place, the bank of the channel of the high floodplain, 07/31/1996.

Carex dichroa (Fr.) V. Krecz. x C. cryptocarpa C.A. Mey. Assembled twice. Pebbles, ducts in the floodplain, bushes of floodplain terraces.

The upper course of the river. Chiranay: p. Mayolgykonvaam in the middle course, the channel in the floodplain, 07/06/1995, The upper course of the river. Nygchekveem: mouth of the river. Ichguigeynei, floodplain terrace, mixed shrubs, 06/30/1996.

Carex eleusinoides Turcz. ex Kunth. - sedge elevzinovidnaya. Everywhere. Sporadically. Floodplain pebbles. The view in our region is associated with icy valleys. Alaskan-East Asian subalpine species of pebbles. From the northern part of the Koryak Upland is known only from the vicinity of the village. Beringovsky. Our plants have roots without green pubescence, the lower spikelet is 1-2 cm apart, the floral scales are already shorter than the sacs. Our plants are most often erect and tall. In general, this species in the region is quite rare and is associated with the icy valleys of rivers and streams.

Upper river Khatyrka: mouth of the creek. May, pebble in the floodplain of the river, 07/12/1993, pebble stream. Maisky, 07/07/1993, River basin. Tamvatvaam: confluence of the Tundra and Tumgytveem rivers, floodplain pebble, 07/20/1994, icy valley in the lower reaches of the river. Tundra, 07.23.1994, the middle course of the river. Tumgystveem, the icy part of the floodplain, gravel, 07/17/1994, Upper course of the river. Chiranay: upper river. Mayolgykonvaam, icy section, 07/17/1995, the mouth of the river. Mayolgykonvaam, high floodplain, willows along the canal, 07/07/1995, The middle course of the river. Chiranay: mouth of the river. Openwork, floodplain pebble, 08/05/1995, Upper river. Nygchekveem: mouth of the river. Ichguigeynei, dry canals on the floodplain terrace, 06/30/1996, the vicinity of lake. Smooth, silty duct in the floodplain, 1.07.1996, Surroundings of Lake Mainz: west coast against Tumanny Island, icy section of the lake terrace, 07.20.1996.

Carex fuscidula V. Krecz. ex Egor. - Sedge is brownish. Everywhere. Not rarely. Tundra, thickets of cedar dwarf, on loamy spots. Siberian-American hypoarcto-montan tundra species. This is essentially a subspecies. C. capillaris L., characterized by small size (medium) and slightly darker spikelets.

Upper riverKhatyrka: intermountain hollow near Mount Evlanyr, high hillocks on the terrace - ledge, 07/20/1993, Tamvatney mountains: Mount Tumannaya, saddle, tundra with dwarf curtains, 07/27/1994, River basin. Tamvatvaam: p. Tundra at the mouth, plume of a hill, forbs and shrubs of tundra, 07.24.1994, in the same place, floodplain terrace, glades in the thickets of cedar dwarf, 07.24.1994, Upper course of the river. Chiranay: mouth of the river. Elgiveem, plume of Mount Chelka, sedge tundra, 07/19/1995, Middle course of the river. Chiranay: mouth of the river. A winding, moraine ridge, loamy spots in the hummock tundra, among thickets of cedar dwarf, 2.07.1995, Sources of the river. Nygchekvei, Koryak Range: the wall of the car of the southern exposure over the glacier at the head of the stream. Nakhodka, 07/07/1996, Upper course of the river. Nygchekveem: the vicinity of Lake. Smooth, shrubby tundra on the floodplain terrace, 06/28/1996.

Carex glacialis Mackenz. - Sedge ice. Everywhere in the mountains. Seldom. Curtain gravelly tundra, nival niches. Circumpolar arctoalpine mountain view. From the central part of the Koryak Upland, one location is known in the area of ​​Mount Ledyanaya. For the northern part of the Koryak Upland is given for the first time. The nearest locations are along the Anadyr and Penzhina rivers. For Kamchatka is not indicated.

River basin Tamvatvaam: the middle course of the river. Tumgytveem, massif of Mount Yagelnaya, kurtin tundra along the ridge, 07/17/1994, Upper course of the river. Chiranay: p. Mayolgykonvaam, nival spots on the first floodplain terrace under the ledge of the high fluvioglacial terrace, 07.17.1995, Sources of the river. Nygchekvei, Koryak ridge: top (ridge) of the mountain between the stream. Rotten and Nakhodka, clumpy gravelly tundra, July 9, 1996.

Carex globularis - Round sedge

Carex glareosa Wahl. - Sedge pebble. Everywhere. Seldom. Sporadically. Ice valleys, lichen heaths of terraces, meadows and nival tundra in hollows, hollows and on ledges of moraines and other fluvioglacial deposits. Circumpolar littoral-arctic species. News for the northern part of the Koryak Highlands. Plants assigned to this species differ from C. tripartita All. only by the fact that male flowers are present only at the base of the upper spikelet, lower spikelets only with female flowers.

Upper river Khatyrka: ruch. Maysky, swamp along a stream, 06/17/1993, river floodplain, pebble, 07/12/1993, thermokarst depressions on the fluvioglacial terrace, 07/12/1993, in the same place, sedge wastelands in depressions, 07/11/1993, River basin. Tamvatvaam: the lower reaches of the river. Tundra, icy site in the floodplain, a grassy meadow under the ledge of a high terrace, 07.23.1994, the mouth of the river. Tundra, lichen wasteland on the floodplain terrace, 07.24.1994, Upper course of the river. Chiranay: mouth of the river. Mayolgykonvaam, nival spots under the ledge of the high fluvioglacial terrace, 07.17.1995, ibid., Mixed grass willows under the ledge, 07.07.1995, Upper course of the river. Nygchekveem: the vicinity of Lake. Smooth, decay on Mount Tumble, nival hollow, 06.26.1996.

Carex globularis L. - Round sedge. Everywhere in the zone of boreal dwarfs. Very common. Tundra, thickets of elfin trees, raw yerniki. Eurasian hypoarctoboreal forest-bog species. In some individuals, the veins on the sacs are poorly distinguishable, the inflorescence is quite compact. These individuals can be defined and how C. vanheurckii Mueller.

Upper river Khatyrka: intermountain hollow near Mount Evlanyr, dry kobresia mounds on the watershed, 07/20/1993, River basin. Koyverelyan: mouth of the river. Koyverelyan, third floodplain terrace, kochkarnaya tundra, 07/29/1993, ibid., Second floodplain terrace, 07/29/1993, River basin. Tamvatvaam: the middle course of the river. Tumgytvayem, mountain trail, erniki, 07/14/1994, Upper river. Chiranay: upper river. Dry, horsetail and shrub tundra on the mountain plume, 07/11/1995, plume of the mountain of the Chirinaisky mountains, horsetail dwarfs, 07/24/1995, The middle course of the river. Chiranay: the vicinity of Lake Good, the slope of the slope, kochkarnaya tundra, 1.08.1995, p. The winding, hummocky tundra with bushes of cedar dwarf on the top of the hill, 2.07.1995, around Lake. Good, hummocky tundra with dwarf pine bushes on a high terrace, 07/30/1995, Upper river.Nygchekveem: against the mountain of George (turn), the sources of the stream. Sentinel, raw yerniks, 07.26.1996, The middle course of the river. Nygchekveem: in the range of the mountain Nygchek - Lake. Eternal, moraine on the right bank of the river, thickets of elfin trees, 07/31/1996, surroundings of Mount Nygchek, moraine on the right bank of the river, thickets of yernik and elfin trees, 08/05/1996.

Carex gynocrates Wormsk. - The sedge is feminine. Seldom. Willows on streams, shrubby tundra on terraces. American-Asian boreal-hypoarctic tundra-bog species. News for the northern part of the Koryak Highlands.

The middle and lower reaches of the Chyrynai river: at the trigopunkt 78 m, shrubby tuberous tundra on the first floodplain terrace of the river, 07/27/1995, Chyrynai mountains: opposite Mount George, turn of the river. Nygchekveem, willows from Salix alaxensis along the brook on the floodplain terrace, 07.24.1996.

Carex holostoma dre. - Sedge one-handed. Collected once. Circumpolar arctic view. News for the Koryak Highlands as a whole.

The upper course of the Chyrynai river: the mouth of the river. Mayolgykonvaam, high (9 m) fluvioglacial terrace, ernik tundra, description 14, July 6, 1995.

Carex juncella (Fries) Th. Fries = C. wiluica Meinsh. - Sedge Sitnik. Everywhere. Not rarely. Shrubs floodplains, shrub tundra terraces and moraine dips. Eurasian boreal-hypoarctic forest-tundra-bog species. For the Koryak Uplands as a whole, is given for the first time. In Chukotka is not known. Many locations in the Magadan region.

Upper river Khatyrka: mouth of the creek. Maisky, thermokarst depressions on the fluvioglacial terrace, 07/12/1993, Tamvatney mountains: low floodplain terrace of the river. Tamvatvaam, shrub tundra, 07/29/1994, River basin. Tamvatvaam: the middle course of the river. Tumgytvey, shrubs of the middle floodplain, 07/16/1994, Upper river. Chiranay: mouth of the river. Mayolgykonvaam, high shrubs of high floodplain, along the canal, 07/07/1995, The middle course of the river. Chiranay: at the trigger point 78 m, tuberous shrub tundra on the terrace, 07.28.1995.

Carex juncella (Fries) Th. Fries ssp. minuta (Fran.) Galanin comb. prov. - Sedge Vilyuyskaya shallow. Seldom. The bottoms of drained moraine lakes. Nearest location C. minuta Fran. - in the middle course of the river. Omolon and on the river. Mal Anyui.

The upper course of the river. Nygchekveem: against Mount Janranai, a deflated lake on the moraine, 07/19/1996.

Carex kreczetoviczii Egorova - Osoka Krechetovicha. Collected once. East Siberian boreal erosiophil species.

Upper river Khatyrka: intermountain depression, the Vililyumeykuul river at the head, willow on the first floodplain terrace, 07.24.1993.

Carex jacutica V. Krecz. - Sedge Yakut. Everywhere in the zone of boreal dwarfs. Pretty rare. Swamps, meadows, willows along the banks of lakes and rivers. East Siberian boreal floodplain-bog species. The closest locations are limestones in the Kolyma Highlands, the upper reaches of the Khatyrka River, the environs of the city of Anadyr, pos. Markovo.

Upper river Khatyrka: ruch. Maisky, an ancient terrace, along a stream, 07/16/1993, in the same place, a swamp along a stream under a mountain trail on a fluvioglacial terrace, 07/17/1993, Basin of the Tamvatvaam river: r. Tundra near its mouth, low floodplain with gullies, willows, 07/25/1994, Upper course of the Nygchevei river: against Mount Janranai, on the shore of Lake. Lowland, 07/19/1996.

Carex lachenalii Schkuhr. - Sedge of Lachenal. Not rarely. Sedge-mixed grass nival tundra, stream banks (Khoreva, 2016). Circumpolar arctoalpine meadow-bog species.

Carex lapponica O. Lang. - Sedge Lapland. Everywhere in the zone of boreal dwarfs. Pebbles, sedge meadows, swamps, bottoms of drained moraine lakes. Circumpolar arctic view of riverine ecotopes and swamps. Soddy plants (var. compacta Galanin) with wide leaves (up to 2 mm) dodging towards C. cinerea Pall., Possibly hybrids with this species.

Upper river Khatyrka: thermokarst depressions on a high terrace of the river, 07/12/1993, Basin of the Tamvatvaam river: watershed of the Tumgytveem and Razliv rivers, sedge-mixed grass swamp, 07/19/1994,
Tamvatney mountains: r. Tamvatvaam, middle floodplain, old pebble, 07/25/1994, Middle and lower reaches of the Chirinai river: the vicinity of Lake. Good, along the edge of a tundra lake, 07/31/1995, in the same place, a flat lake in the tundra, 08/08/1995, the middle and lower reaches of the Velikaya River: Peschany island against a transshipment of oil workers, potholes among a swamp on a floodplain terrace, 08/17/1995 , transshipment of oil industry workers, swamp in the lowland tundra, 08/09/1993, Upper course of the Nygchekvei river: against Mount Janranai, shore of lakeLowland, sedge meadow, 07/19/1996, Middle course of the Nygchekvei river: right bank to the Nygchek mountain, moraine, sedge swamp in the thermokarst failure, 07/31/1996.

Carex ledebouriana C.A. Mey. ex Trev. - Sedge of Ledebour. Seldom. Slope shrub tundra. Asian arctoalpine mountain meadow view.

Surroundings of the city of Anadyr: the slope of Mount Dionysius, description 5, 2.07.1993.

Carex leporina L. ssp. korjakensis Galanin nom. prov. - Sedge rabbit Koryak. Collected once. Our plants are similar to the description C. haydeniana Olney from Sev. America (Hithcock, Cronquist, 1973), but differ in the absence of veins on the sacs and the presence of denticles in their upper part. From C. pallida C.A. Mey. differ in that all spikelets in a dense capitate ovate inflorescence are gynecandric, i.e. with pistil flowers at the top. In addition, does not form turf. Our plants have sacs without noticeable veins, the nose of the sac is distinctly jagged. Plant height 25-30 cm, stems almost half leafy. Leaves 3-4 mm wide. Rhizome plants. Inflorescences 1x 1.5 cm. Bract leaf scaly. Scales of pistil flowers are light brown, equal or longer than the sacs. Sepals of stamen flowers white-edged along the edge.

Upper river Khatyrka: pebble on the old river terrace, shrubs on lichen wasteland, 07/16/1993.

Carex limosa L. - Moss sedge. Seldom. Sedge marshes in thermokarst moraine dips. Circumpolar hypoarctoboreal bog species. Meets with C. magellanica Lam., From which it differs by clear veins on the sacs and their number, shorter leaves. Fees are known from the hall. Corfu, the basins of the Penzhina and Anadyr rivers.

The middle course of the Nygchekvei river: in the range of the Nygchek mountain and lake. The eternal, right bank, sedge swamp in the thermokarst failure on the moraine, 07/31/1996.

Carex lugens H.T. Holm. - Sedge is brilliant. Everywhere. Usually. Meadows, willows, alders, moist tundra, swamps. Alaskan-East Asian arctic char. In our region, typical C. lugens is very rare. Most fees still apply to C. soczavaeava Gorodk., Which can be considered a subspecies C. lugens. Often there are forms that evade C. ensifolia Turcz. ex Krecz. and C. caespitosa L. s.l. - probably hybrids with these species.

Upper river Khatyrka: thermokarst subsidence on the fluvioglacial terrace at the mouth of the creek. Maisky, 07/12/1993, lichen tundra with thickets of cedar dwarf on a moraine terrace, 07/16/1993, intermountain hollow near Mount Evlanir, high floodplain, 07/20/1993, River basin. Tamvatvaam: the Tundra River in the lower reaches, a floodplain terrace, a tuberous tundra with an ernik, 07/25/1994, p. Tundra at the mouth, log on the hillside, sedge-shrub tundra, 07.27.1994, Tamvatney mountains: mountain pass, sedge mixed grass tundra near the watershed lakes, 07.26.1994, Upper Chyrynai river: against the Chirinai mountains, meadow on the high slope terraces, precipice, 07.24.1995, the Axial part of the Koryak ridge, the sources of the river. Find: raw meadow and swamp on a mountain plume, 07/06/1996, a moss-grass swamp on a slope plume, 07/06/1996, Upper river. Nygchekveem: Lake Smooth, hollow of a drain at the lake, willow from Salix pulchra, 06/29/1996, against Mount Janranai, moraine hillocks on a slope in Lake. Mainits, alder-houses, 07.20.1996, Surroundings of the city of Anadyr: a plume of the slope of Mount Dionysia, raw hummock tundra, 2.07.1993.

Carex mackenziei V. Krecz. - Sedge Mackenzie. Rarely on marching ecotopes along the coast of the Anadyr Gulf. Tame tundra. Amphibipolar seaside species.

Surroundings of the city of Anadyr: south of Mount Dionysia, along a watercourse in the Second River area, 06/20/1993.

Carex melanocarpa Chamisso ex Trautvetter - Sedge Aronia. Everywhere in the zone of boreal dwarfs. Curtain, shrub tundra, shrub tundra with thickets of cedar dwarf. Pretty common. Less commonly in the nival tundra in the adjacent areas. Siberian hypoarcto-montane meadow-tundra species. Some individuals form very loose turfs with powerful leaves 1/3 shorter than generative shoots.

Upper river Khatyrka: moraine subsidence near the mouth of the stream. Maisky, 07/07/1993, lichen tundra with curtains of cedar dwarf, high fluvioglacial terrace, 07/16/1993, Basin of the Tamvatvaam river: the middle course of the river.Tumgytveem, Yagelnaya mountain, kurtina tundra on the slope, 07/17/1994, Tamvatney mountains: Shamanya mountain, log in the lower part of the northern slope, herb-shrub tundra, 07/12/1994, ibid., Rocky-gravelly tundra on the slope, 07/12/1994, high terrace of the Tamvatvaam river, shrub tundra with cedar dune curtains, 07/30/1994, Martian mountain, gravel talus on the slope, clump tundra, 07/26/1994, Upper Chyrynai river: Ugryumaya mountain, tundra in the saddle with loamy spots, 07.07. 1995, the top of Mount Ugryumaya, the curtain tundra, 07/13/1995, Mount Yyl, the curtain ivershinnaya tundra, 08.07.1995, axial part of the Koryak Range, the origins of p. Find: wall of the car of the southern exposure above the glacier, 07/07/1996, nival herb-dryad tundra on the slope, 07/05/1996, lamb foreheads in the puncture system of the stream. Vostochny, 07/10/1996, Upper course of the river. Nygchekveem: the vicinity of Lake. Smooth, shrubby tundra on the floodplain terrace, 06/28/1996, ibid., Basin on the moraine ridge, 06/28/1996, The middle course of the river. Nygchekveem: the tract Maino-Gytkino, conflagration on the fluvioglacial terrace, 08/09/1996.

Carex membranacea hook. - Sedge webbed. Everywhere. Usually. Floodplain ecotopes. By streams can dominate. Less commonly in the tuberous tundra. Alaskan-East Siberian arctic boreal floodplain meadow species. Sacks and scales are dark-colored - from dark brown to almost black, the base of the stems and rhizomes are covered with purple scales (to dark cherry). The upper female spikelet at the top sometimes has male flowers.

Upper river Khatyrka: thermokarst subsidence on the fluvioglacial terrace near the mouth of the brook. Maisky, 07/12/1993, in the same place, alder on a terrace ledge, 07/12/1993, a swamp along a brook on a high terrace, 07/16/1993, intermountain hollow near Evlanir hill, dominant on a brook, 07/20/1993, in the same place, a floodplain terrace of the river, 07/20/1993, lake Meingautgytkhyn, on the shore, 07/14/1993, Basin of the Tamvatvaam river: the middle course of the river. Tundra, icy section, 07/21/1994, the middle course of the river. Tumgytvey, shrubs of the middle floodplain, 07/16/1994, p. Tundra at the mouth, flood-plain willow willow, 07/24/1994, Upper course of the Chyrynai river: mouth of the river. Mayolgykonvaam, high fluvioglacial terrace, willows along a dry temporary groove, 07/04/1995, ibid., Willows of a high floodplain, 07/07/1995, Mount Ugryumaya, valley of the river. Vilyuy, shrubs of the first floodplain terrace, 07.15.1995, Middle and lower reaches of the Chyrynai River: the vicinity of Lake Good, alder-willow trees on the 1st floodplain terrace, 07/30/1995, shrubby tuberous tundra on the floodplain terrace, 07/27/1995, Axial part of the Koryak Range, the sources of the river. Find: mixed-moss swamp on a train, 07/06/1996, Upper course of the river. Nygchekveem: near the mouth of the river. Kelmyveem, silted duct in the floodplain, 07/16/1996, the vicinity of lake. Smooth, terraces in the valley brook. Old, 06/26/1996, against the mountain of George (turn), willow along the brook on the terrace, 07/24/1996, against the mountain Yanranay, the shore of the lake. Lowland, 07/19/1996,
The average course of the river. Nygchekveem: mixed grass shrubs of the floodplain terrace in the range of the mountain Nygchek and Lake. Eternal, 07/29/1996, Lake Mainitz region: lakeside terrace on the west bank, icy plot, willow from Salix saxatilis, 20.07.1996.

Carex micropoda C.A. Mey. - Sedge small-footed. Collected once in the area of ​​the village. Beringovsky. Nival meadows along mountain streams. The Far Eastern North American, probably the Bering Arctic species. It is listed in the Red Book of Chukotka (2008) as a rare species.

Carex middendorfii Fr. Schmidt. - Sedge Middendorff. Collected once. Far Eastern boreal bog species.

Basin of the Tamvatvaam River: the middle course of the river Tumgytvey, watershed with the river. Spill, sedge-mixed bog, 07/19/1994.

Carex misandra R. Brown. - Sedge is a man-hating. Everywhere. Not rarely. Different types of wet and moist tundra, especially with elements of nivation. Circumpolar arctic tundra species. Sacks in the upper part along the edges are prickly, elongated, gradually turning into a nose.

Tamvatney mountains: Martian mountain, steep northern slope above the snowfield, scree, 07.26.1994, ibid., Kurtino vegetation on fine-grained scree, 07.27.1994, in the same place, spotted motley grass tundra with stalnik in the lower part of the mountain slope, 07.26.1994, there the loamy-gravelly plume, sedge tundra, 07.26.1994, gravelly slopes of the mountains, spotted tundra, 07.31.1994, Upper course of the river. Chiranay: top of Mount Ugryumaya, spotted tundra on the slope, 07/13/1995, Mount Yyl, kurtinny tundra in the upper part, 07/08/1995,
The axial part of the Koryak ridge, the sources of the Nakhodka river: grass tundra on the floodplain terrace of the stream, 07/07/1996, in the same place, near the glacier in the square, the dryad nival tundra, 07/07/1996, in the same place, lamb foreheads on the watershed between two lakes at the source of the p river . Vaamochka, 07/10/1996, wall of a car of southern exposure over a glacier, 07/07/1996, Upper course of the Nygchevei river: opposite Mount George (turn), raw hummock tundra at the foot of the mountain 450 m, 07.24.1996, Outskirts of Anadyr: slope trail Mount Dionysia, raw hummock tundra, 2.07.1993.

Carex molissima Christ. - Sedge is the softest. Seldom. Tame bogs and thermokarst depressions on fluvioglacial terraces. Siberian boreal forest-bog species. The nearest locations are in the Kolyma Upland and the continental regions of Okhotia. Our samples are well consistent with the description of the species and are quite typical for it. The difference is that some individuals develop one rather than two stamen spikelets.

Upper river Khatyrka: mouth of the creek. Maisky, thermokarst depressions on the fluvioglacial terrace, 07/12/1993, in the same place, a swamp along a stream on a high terrace, 07/16/1993.

Carex nesophila H.T. Holm. - Sedge is island-loving. Seldom. In the hollows of high floodplain terraces, in the bushes of the floodplain. Beringian arctic tundra view. Our locations are the southernmost in the range of the species. In our area, typical individuals of this species are extremely rare. In all likelihood, this species should be considered a species. C. podocarpa ssp. podocarpa.

Upper river Khatyrka: depressions on a high terrace at the mouth of the creek. Maisky, 07/12/1993, Basin of the Tamvatvaam River: the middle course of the river. Tumgytveem, high floodplain terrace, in the hollows, 07/14/1994, Upper course of the Chyrynai river: mouth of the Mayolgykonvaam, high floodplain, high shrubs, 07/07/1995.

Carex norvegica Retz. Retz. ssp. angarae (Steudel) Galanin comb. nov. - Sedge Norwegian. Everywhere. Usually. By waterways, gravels of floodplains and floodplain terraces. North American-Eurasian boreal meadow species. Our plants can have black, dark brown, brown, and light brown scales. Pouches are most often veined; leaves are often half shorter than the stems. Forms turfs, sometimes small rather dense bumps.

Upper river Khatyrka: floodplain terrace Maisky, 07/17/1993, intermountain depression near the Evlanir hill, high floodplain, 07/20/1993, intermountain depression, a lake in the tundra, along the coast, 07/20/1993, Tamvatvaam river basin: the middle course of the river. Tundra, icy valley, 07/23/1994, in the same place, floodplain Chosenia forest, description 22, 07/21/1994, the lower reaches of the river. Tundra, icy section, 07/23/1994, Upper course of the Chyrynai river: the upper reaches of the river. Mayolgykonvaam, icy section, 07/17/1995, Ugryumaya mountain area, floodplain terrace of the river. Vilyui, mixed shrubs, 07.15.1995, Upper course of the Nygchekvei river: 3 km above the mouth of the river. Ichguigeynei, a chozen forest in the floodplain, 07/12/1996, against Mount Yanranai, a fluvioglacial terrace, an all-terrain road, 07/19/1996, against Mount George (turn), mixed willows on a rivulet on a terrace, 07/24/1996, Lake District. Mainz: the western shore of the lake, the icy section of the lakeside terrace, 07.20.1996.

Carex obtusata - sedge blunt

Carex obtusata Liljebl. - sedge blunt. Seldom. Dry gravelly tundra, steppe areas, alder on rocky scree. Disjunctive-circumpolar plurizonal mountain-steppe species. Our plants shy away from C. gynocrates Wormskjold. For the Koryak Uplands is given for the first time, the closest location - north of the village. Coal Mine on the banks of the Anadyr Estuary. In Kamchatka is not known.

Tamvatney mountains: Tumannaya mountain, a steep northern slope, alder clumps on talus, 07/27/1994, dry gravelly plume with dry mounds, steppe tundra, 07/27/1994, top of the mountain, dry gravelly shrubby tundra, 07/31/1994.

Carex pallida C.A. Mey. - sedge pale

Everywhere in the zone of boreal dwarfs. Floodplain pebbles, floodplain and floodplain meadows, shrubs, alders. East Siberian-Far Eastern boreal-montane meadow-forest species. For the Koryak Upland, it is given only for Korfa Bay, but it is not infrequent in the Penzhina basin and on the river. Anadyr. In our plants, the sacs are bare, strongly serrated along the edge.

River basin Koyverelyan: high floodplain in the middle reaches of the river. Koyverelyan, poplar forest, 07.26.1993, Middle and lower reaches of the river. Great: 5 km below the mouth of the river. Chyrynay, high floodplain on the left bank, meadows, willows-copses, 08/12/1993, the mouth of the river. Koyverelyan, alder forest in the floodplain, 07/29/1993, Tamvatney mountains: low floodplain terrace of the river.Tamvatvaam, shrub tundra, 07/29/1994, alder-willow forest in the floodplain of the river. Tamvatvaam, 07/27/1994, The middle course of the river. Chiranay: okr. Lake Good, alder-meadow, 1st floodplain terrace, 07/30/1995, p. Openwork, floodplain drying alder, 08/05/1995, Upper course of the river. Nygchekvei: opposite Mount George (turn), a bush on a floodplain pebble, 07/27/1996 (not seen above the river Nygchevei), the average course of the river. Nygchekveem: in the range of the mountain Nygchek - lake. Eternal, mixed shrubs on the first floodplain terrace, 07/31/1996.

Carex paupercula Michx. - Sedge depleted. Seldom. Sedge marshes in thermokarst moraine dips. Circumbipolar boreal bog species. The nearest species locations are the hall. Korfa, Kamchatka and Okhotia from the Pit to Tauya, South Okhotia and Amur.

The middle course of the Nygchekvei river: in the range of the Nygchek mountain and lake. The eternal sedge swamp in the thermokarst failure on the moraine, 07/31/1996.

Carex petricosa Dewey. - Sedge rocky

It is found in hyperbasites in our area. Rarely, we collected it only once. The Beringian species, indicated only for the east of the Chukchi Peninsula and in the Kolyma Upland as a calcephilus. News for the flora of the Koryak Upland, the closest locations are in the east of the Chukchi Peninsula and in the Kolyma Upland. Calcephilus.

Chiranay mountains, pike meadow tundra on the mountain terrace, 07/22/1995.

Carex podocarpa R. Brown ssp. koraginensis Meinsh. comb. prov. - Sedge Nogorozhnaya Coraginskaya (meadow form C. podocarpa). In the flora of neighboring regions is indicated as C. koraginensis Meinsh. Everywhere. Often. The most common type of sedge in the region. Meadows, tundra and dwarf lands of all types of habitats, rocky scree, floodplain pebbles and rocks. East Siberian, mainly North-Pacific meadow-tundra nival species. Bags with spines along the upper edge, part of female spikelets almost always with male flowers, scales at the base from brown to chestnut, split into long fibers. Bracts vary greatly in width - wider, equal or narrower than sacs - and in color. The stems are rough.

Upper river Khatyrka: ruch. Maisky, high terrace, ravine with a stream, 07.16.1993, p. Meingutvaam, alder on the slope, 07/14/1993, floodplain Maisky, 07/07/1993, top of the mountain, in placers, 07/08/1993, brook. Maisky, high river terrace, thermokarst hollow, 07/12/1993, Tamvatney mountains: mountain hollow in the lower part, alder, 07/27/1994, mountain pass, mixed sedge tundra in the through valley, 07/27/1994, log in the lower part of the northern slope , forbs and shrubs tundra, 07/12/1994, lower part of the Martian slope, mottled spotted tundra with dwarf, 07/26/1994, sedge tundra on a plume, 07/26/1994, River basin. Tamvatvaam: Mount Yagelnaya in the middle reaches of the river. Tumgytveem, glade in the dwarf thickets on the floodplain terrace, 07.17.1994, Upper course of the river. Chiranay: mouth of the river. Mayolgykonvaam, high moraine terrace, July 4, 1995, in the same place, dry duct on the terrace, July 4, 1995, in the same place, dry bushes of the first floodplain terrace, July 7, 1995, Mount Chelka, sedge tundra, July 19, 1995, Middle course of the river. Chiranay: mouth of the river. Openwork, floodplain, willows and alder, 08/05/1995, ibid., Pebbles of the floodplain with willows, 5/08/1995, okr. Lake Good, meadow on the ledge of a high (13 m) terrace, 07/30/1995, in the same place, meadow on the ledge of a high (6 m) terrace, 08/08/1995, Sources of the river. Nygchekvei, Koryaksky ridge: nival forbs and dryad tundra on the slope, 07/05/1996, upper stream. Nakhodka, raw mixed tundra on a slope plume, 07/06/1996, Upper course of the river. Nygchekveem: mouth of the river. Ichguigeynei, floodplain gravel, 30.06.1996, ibid., Mixed shrubs on the floodplain terrace, 06/30/1996, okr. Lake Smooth, terraca valley creek. Old, 26.06.1996, in the same place, a floodplain terrace, shrubby tundra, 28.06.1996, in the same place, floodplain pebbles, 07/03/1996, in the same place, nival decay on Mount Osypnaya, 06/26/1996, against Mount George (turn) , cliffs of the hyperbasitic mountain 284 m, along a stream in the nival hollow, 07.24.1996, Middle course of the river. Nygchekveem: the shore of a moraine lake at the foot of Mount Nychek, 08/08/1996, the bank of the river. Silvery - a hillock-outlier of a high terrace, 07/29/1996, in the alignment of Mount Nychek - lake. Eternal, ledge of a moraine terrace with thickets of cedar dwarf, 07/29/1996, Surroundings of the lake. Mainz: western shore of the lake, alder forest in the upper part of the slope, 07.20.1996.

Carex podocarpa R. Br. ssp. podocarpa - Sedge Nasal (nival form of C. podocarpa). Everywhere, but not as often as the Coragin subspecies. Sporadically. Nival forbs and shrubs of the tundra.East Asian-North American boreal species of wide ecological amplitude. Known collection from the village. Beringovsky. Plants up to 20-30 cm tall, female spikelets ovate, scales and sacs almost black, sacs practically without spines, smooth stems. Female spikelets without male flowers.

Tamvatney mountains: Mount Shamanya, a log in the lower part of the sowing slope. expositions, forbs and shrubs of the tundra, 07/12/1994, Sources of the river. Nygchekvei, Koryak Range: eastern slope, nival tundra, 07/06/1996, herb-dryad tundra on the slope, 07/05/1996, the upper part of the slope zap. expositions, mixed grass tundra, 07/09/1996, mixed grass tundra at the bottom of the Kara at Lake. Lonely, 07/05/1996, snowy meadows in the kar system of the brook. East, 07/10/1996.

Carex podocarpa R. Br. ssp. pulchra Galanin nom. prov. - Sedge beautiful breast (mottled form in hyperbasites). Everywhere. Usually, but only in areas of hyperbasite distribution. Pebbles, meadows and tundra. It features light colored bags and narrow and short brown scales. The plants are taller, the stems are 2-3 times larger than the root leaves. The bags are densely covered with spines. Most similar to C. macrogyna Turcz. ex Steud., from which it is clearly different in that the upper spikelet is always male without female flowers (a sign of the group C. podocatrpa).

Tamvatney mountains: meadow under the terrace ledge, 07/30/1994, Upper river. Chyrynai: against the Chyrynai mountains, a cliff on the slope of a high terrace, meadow, 07.24.1995, The middle course of the river. Chiranay: near the mouth of the river. Openwork, the first floodplain terrace, the bank of the canal with ravines, mixed shrubs, 6.08.1995, Upper course of the river. Nygchekveem: against Mount George (turn), hyperbasitic mountain 284 m, raw tundra with cedar dwarf on the slope, 07.24.1996, Upper course of the river. Nygchekveem: okr. Lake Smooth, floodplain pebble, 07/03/1996.

Carex rariflora (Wahlb.) Smith. - Sedge rarely flowered. Everywhere. Not rarely. Swamps, tuberous shrubby tundra, icy sections of valleys. Circumpolar arctic bog species. News for the northern part of the Koryak Highlands. The nose of the bags is missing. Sacks with obscure veins. The scales are slightly shorter than the mature sacs and are already equal and slightly longer and wider; the color of the scales varies from light brown to black.

Upper river Khatyrka: mouth of the creek. Maisky, thermokarst depressions on a high moraine terrace, 07/12/1993, in the same place, alder on a terrace ledge, 07/12/1993, in the same place, eutrophic swamp on a high terrace, 07/16/1993, River basin. Tamvatvaam: watershed of the Tumgytveem and Razliv rivers, sedge-mixed grass swamp, 07/19/1994, Upper course of the river. Chiranay: swamp in the valley of the river. Vilyuy, 07.15.1995, eutrophic tuberous bog with pits in the valley of the stream. Sukhoi, 07/11/1995, the upper river. Mayolgykonvaam, the icy section of the valley, 07/17/1995, The middle course of the river. Chiranay: at the trigger point 78 m, tuberous shrubby tundra, 07.28.1995, Sources of the river. Nygchekveem, Koryak Range: a swamp on a terrace at the mouth of the creek. Nakhodka, 07/11/1996, a moss-grass swamp on a plume of a slope of eastern exposure, 07/06/1996, Upper course of the river. Nygchekveem: mouth of the river. Kelmyveem, a swamp in a river valley, 07/16/1996, against Mount George (turn), sedge swamp on a moraine, 07/24/1996, okr. Lake Smooth, pits along the lake, 06/29/1996, The middle course of the river. Nygchekveem: in the range of the mountain Nygchek and Lake. The eternal sedge swamp on the floodplain terrace, 07.29.1996, okr. Nychek mountains, Lake Lonely Owl, on a moraine along the lake, 08/05/1996, Surroundings of the lake. Mainz: the western shore of the lake against Tumanny Island, the key swamp on the moraine terrace, 07/20/1996, Station "Oneman": the hummock tundra on the high root terrace - uval, 09/03/1993.

Carex rhynchophysa - sedge swelling

Carex redowskiana C.A. Mey. - Sedge Redowski. Everywhere in the zone of boreal elfin and southern hypoarctic tundra. Not rarely. Swamps, willows along streams. East European-Siberian arctoboreal taiga-bog (?) Species. Our plants should rather be attributed to C. gynocrates Wormsk. The news for the Koryak Upland as a whole is not noted in Kamchatka either.

Upper river Khatyrka: Khatyr intermountain depression, p.Vilyumeikuul in the headwaters, 1st floodplain terrace, willow, 07.24.1993, high terrace brook. Maisky, thermokarst depressions, 07/12/1993, Upper river. Chiranay: swamp at the mouth of the river. Vilyuy near Ugryumaya Mountain, 07/15/1995, The middle course of the river. Chiranay: okr. Lake Kind, drying shack in the tundra, 07/31/1995, Upper river. Nygchekveem: near the mouth of the river. Kelmivem, swamp in the valley, 07/16/1996, against Mount George (turn), willows from S. alaxensis along the brook on the terrace, 07.24.1996, the Middle course of the river. Nygchekveem: near the mountain Nygchek, sedge swamp near the lake on the terminal moraine, 08/03/1996.

Carex rhynchophysa C.A. Mey. - sedge swelling. Not rarely, but sporadically. On the banks of ponds, floodplain meadows. Almost circumpolar boreal bog species. News for the northern part of the Koryak Highlands.

Upper river Khatyrka: high terrace at the mouth of the stream. Maisky, eutrophic swamp along a stream, 07.16.1993, ibid., Thermokarst depressions on a high terrace, 07.12.1993, Middle and lower reaches of the river. Velikaya: Peschany island against a transshipment of oil workers, a 4 m high terrace, a reed meadow along the canal, 08/17/1995, the mouth of the river. Koyverelyan, silty duct in the floodplain, alder forest, 07.29.1993, Velikaya River, 6 km above the weather station, meadow along the duct, 08.15.1995.

Сarex rostrata - Nose sedge

Сarex rostrata Stokes. - Nose sedge. Sporadically. Swamps. Circumpolar hypoarctoboreal bog species. News for the northern part of the Koryak Highlands.

Middle and lower river. The Great: against Peschany Island at the oil depot, duct on a high (4 m) terrace, 08/17/1995, Middle course of the river. Chiranay: okr. Lake Good, meadow at the bottom of a flat lake in the tundra, 1.08.1995, ibid., Swamp along the edge of the lake, 07/31/1995, The middle course of the river. Nygchekveem: terrace by the river. Silver, drained lake on the moraine, 07/29/1996, in the same place, in the lake on the floodplain terrace of the river. Nygchekveem in the area. Lake Eternal, 07/29/1996, moraine under the mountain Nygchek, lake. Lonely Owl, in the water and along the shore, 08/05/1996.

Carex rotundata Wahl. var. tipica - round sedge. Seldom. Raw tundra, swamps, shores of lakes. Almost circumpolar hypoarctic bog species.

Surroundings of the city of Anadyr: Mount Dionysia, slope plume, raw tundra, 2.07.1993, River basin. Tamvatvaam: the source of the river. Spill, shore of the lake on the moraine, 07/19/1994, The middle course of the river. Chiranay: okr. Lake Good, swamp along the edge of the lake in the tundra, 07/31/1995.

Carex rotundata Wahl. var. longida Galanin nom. prov. - Sedge roundish elongated

Everywhere in the zone of boreal elfin and southern hypoarctic tundra. Usually. Swamps, moist tundra, ridge-ridge systems. Our plants are distinguished by a greater height (40–45 cm) and larger cylindrical pistillate spikelets.

Middle and lower river. The Great: transshipment of oil industry workers, pit in the tundra, 08/10/1993, in the same place, a swamp in the lowlands, 08/08/1993, 13 km above the mouth of the Tamvatvaam, raw tundra on a high terrace, swamp, 2.08.1993, Upper river. Chiranay: ruch. Dry, eutrophic tuberous swamp in the valley, 07/11/1995, Middle course of the river. Chiranay: okr. Lake Good ridge-ridge structure in the Kochkar tundra, 1.08.1995, Middle course of the river. Nygchekveem: okr. Lake Eternal, moraine, sedge swamp in the thermokarst failure, 07/31/1996, Outskirts of the city of Anadyr: Mount Dionysia, a plume, raw tundra, 2.07.1993.

Carex rotundata Wahl. x C. membranacea hook. Collected once. Flat narrow leaves, slightly colored base of the stems (with a reddish-brown tint). Flat narrow leaves, slightly colored base of the stems (with a reddish-brown tint). The rest is the same as C. rotundata var. longida.

The upper course of the river. Chiranay: ruch. Dry, eutrophic tuberous bog with pits in the valley, 08/11/1995.

Carex rotundata Wahl. x C. rostrata L. Collected twice. The banks of moraine lakes, the banks of streams along sedge marshes.

River basin Tamvatvaam: the swampy shore of a moraine lake on the watershed of the Tumgytveem and Razliv rivers, 07/19/1994, Upper course of the river. Nygchekveem: against Mount George (turn), a stream along a sedge swamp, 07.24.1996.

Carex rupestris Allioni - Sedge Rocky. Everywhere in the zone of boreal elfin and southern hypoarctic tundra. Pretty rare. Dry spotted and kurtin gravelly tundra on the tops and in the upper part of the slopes. Circumpolar arctoalpine mountain-tundra species. For the northern part of the Koryak Upland, the report “Vascular Plants of the ADD” is not given, but is common in Chukotka and in the southern part of the Koryak Upland.

Tamvatney mountains: top of the Martian mountain (634 m), spotted dryad tundra, 07.26.1994, high terrace of the river. Tavatvaam, shrub tundra with dwarf bushes, 07/30/1994, Upper river.Chiranay: top of Ugryumaya mountain, gravelly mudflow tundra, 07/13/1995, Yyl mountain, spotted and gravelly tundra in the upper part of the mountain, 07/08/1995.

Сarex sabulosa Turcz. ex Kunth = C. melanantha var. sabulosa Kuk - Sand sedge. Collected twice, but not at the place of collection. Swamps, shores of silty channels. Asian hypoarctoboreal type of river sand. From C. glareosa Wahlb. differs in the absence of veins on sacs, does not form a sod - there are long rhizomes. Sagittarius three. For the North-East of Russia it was not previously noted. In the Arctic flora of the USSR, the closest locations are indicated in the lower reaches of the river. Lena, in the upper reaches of the river. Yenisei, in the river basin Vilyuy in Yakutia, in the North. Mongolia and Central Asia. In the summary A.P. Khokhryakova (1985) this species is not given.

Upper river Khatyrka: ruch. Maisky, 3 km above the mouth, a swamp along a stream, 06/17/1993, Upper course of the river. Nygchekveem: okr. Lake Smooth, bank of a muddy channel in a floodplain among willows and alders, 3.07.1996.

Carex sedakowii - Sedoka Sedakova

Carex saxatilis L. ssp. laxa (Trautv.) Kalela = C. procerula V. Krecz. - Sedge stone. Everywhere in the zone of boreal elfin and southern hypoarctic tundra. Usually. Silted coasts, sands, meadows along the channels, floodplain shrubs, bottoms of drained moraine lakes. The Siberian-American hypoarcto-montian subspecies of the arctoboreal meadow-bog species, also common in Europe. For the northern part of the Koryak Upland is given for the first time. Our plants are taller and vary in length along their bracts. The bags and scales are not black, but brown and dark brown, the scales are shorter than the mature bags by almost 1/3 of the length of the latter. The stems and branches of the inflorescence are rough.

Upper river Khatyrka: a watershed plain at the head of the river. Vilyumeikuul, 1st floodplain terrace, willow, 07.24.1993, Middle and lower reaches of the river. Velikaya: 6 km above the weather station, meadow on a silty sandy beach, 08/15/1995, in the same place, meadow along the canal, 08/15/1995, Tamvatney mountains: low floodplain terrace of the river. Tamvatvaam, shrub tundra, 07/29/1994, River basin. Tamvatvaam: high floodplain terrace of the river. Tumgytveem, pit, 07/14/1994, dividing log in the middle reaches of the river. Tumgytveem, sedge bushes, 07/17/1994, Upper river. Chiranay: mouth of the river. Mayolgykonvaam, high floodplain, willows, 07/07/1995, Mount Ugryumaya, floodplain p. Vilyui, willows, 07/15/1995, The middle course of the river. Chiranay: near the mouth of the river. Zhurnaya, sandy river bank Canyonnaya, 08/08/1995, at the trigger point 78 m, shrubby tuberous tundra, 07/28/1995, Upper course of the river. Nygchekveem: against Mount Janranai, a flat lake on the moraine, 07/19/1996, in the same place, the shore of the lake. Lowland, 07/19/1996, against Mount George (turn), a hollow on the terrace, moss-lichen tundra, 07/26/1996, The middle course of the river. Nygchekveem: in the range of the mountain Nygchek and Lake. The eternal sedge swamp along the tundra stream and lake on the first floodplain terrace, 07/29/1996, the bottom of a drained moraine lake near the river. Silver, 07/29/1996.

Carex schmidtii Meinsh. - Sedge Schmidt. Everywhere in the zone of boreal dwarfs. Usually. Floodplain and floodplain meadows, banks of the channel, depressions on moraines. Far Eastern nemoral meadow-bog species. Our plants have a light rather than light brown nose of the pouch, and scales at the base without a pink hue (dark cherry), which reminds C. caespitosa L. Only the well-developed bract leaf differs from the latter.

Upper river Khatyrka: high moraine terrace ruch. Maisky, depressions on the terrace, 07/12/1993, River basin. Koyverelyan: mouth of the river. Koyverelyan, 2nd floodplain terrace, iris meadow, 07/29/1993, Tamvatney mountains: low floodplain terrace of the river. Tamvatvaam, shrub tundra, 07/29/1994, River basin. Tamvatvaam: the middle course of the river. Tumgytveem, depressions on the floodplain terrace, 07/14/1994, Upper river. Chiranay: mouth of the river. Mayolgykonvaam, high terrace, 07/04/1995, The middle course of the river. Chiranay: mouth of the river. Openwork, raw duct in the floodplain alder, 08/05/1995.

Carex sedakowii C.A. Mey. ex Meinsh. - Sedokova Sedakova. Everywhere. Infrequently. Sludge ducts, wet screes under snowfields, loamy spots in the hummock tundra among the elfin thickets. East Siberian alpine-tundra forest-tundra borealmontan species. News for the Koryak Upland and in general the entire ADD, including the Magadan Region. and Kamchatka. Known from the mouth of the river. Penzhins. This is essentially a subspecies. C. capillaris L., characterized by small size and low-flowered spikelets.

Tamvatney mountains: Martian mountain, a steep northern slope above the snowfield, talus, 07.26.1994, Middle course of the river. Chiranay: p.A winding, moraine ridge, loamy spots in the hummock tundra among the dwarf thickets, 2.07.1995, Upper course of the river. Nygchekveem: against the mountain of George (turn), along a stream on the terrace, willows, 07.24.1996, okr. Lake Smooth, silty duct in the floodplain, July 3, 1994.

Carex scirpoidea - Reed sedge

Carex scirpoidea Michaux. - Reed sedge. Everywhere in the zone of boreal elfin and southern hypoarctic tundra. Usually. Forbidden meadows, forbs, forbs, shrubs, and yernik tundra on slopes and moraine ridges. The species is clearly morphologically stable, tends to eutrophic habitats. Most abundant in hyperbasites in moist and normally moistened gravelly-loamy tundra. North Asian-American Arctic mountain tundra species.

Upper river Khatyrka: mouth of the creek. Maisky, a grassy meadow along the bottom of a dry channel on a high river terrace, 07/16/1993, river terrace. Meingutvaam, 07/14/1993, The middle course of the river. Khatyrka: River basin Koyverelyan: Middle and lower reaches of the river. Great: Tamvatney mountains: the lower part of the Martian mountain slope, spotted grass tundra, 07.26.1994, ibid., Sedge tundra on a plume, 07.26.1994, cedar dwarf curtains on a high terrace, 07.30.1994, Mount Shamanya, gravelly-rocky slope, dryad tundra, 07/12/1994, ibid., log in the lower part of the northern slope, herb-shrub tundra, 07/12/1994, River basin. Tamvatvaam: The upper course of the river. Chiranay: mouth of the river. Mayolgykonvaam, loamy spots on a high fluvioglacial terrace, ernik tundra, 07/06/1995, willows along a channel in the middle reaches of the river. Mayolgykonvaam, 07/07/1995, top of Ugryumaya mountain, 07/13/1995, The middle course of the river. Chiranay: Sources of the river. Nygchekvei, Koryaksky ridge: alder forbs on the slope, 07/06/1996, nival forbs and dryad tundra on the slopes, 07/05/1996, Upper river. Nygchekveem: against the mountain of George (turn), the foot of the mountain 284 m, raw kochkarnaya tundra, 07.24.1996, okr. Lake Smooth, moraine ridges, 2.07.1996, ibid., Shrubby tundra on the floodplain terrace, 06.28.1996, against Mount Janranai, flat lake on the moraine, 07.19.1996, Middle course of the river. Nygchekveem: Surroundings of the lake. Mainz: Outskirts of the city of Anadyr: Mount Dionysia, along the watercourse, 2.07.1993, Mount Dionysia, nival tundra on a ledge, 2.07.1993.

Carex soczavaeana Gorodk. - Sedge of Sochava. Everywhere in the zone of boreal elfin and southern hypoarctic tundra. Usually. Raw tundra, swamps, bushes along the hollows. North-East Siberian hypoarctoboreal forest-tundra-bog species. In all likelihood, this is a subspecies. C. lugens, differing only in habit and less compact inflorescence.

Upper river Khatyrka: mouth of the creek. Maisky, thermokarst subsidence on the fluvioglacial terrace, 07/12/1993, ibid., Lichen tundra with dwarf thickets on the terrace, 07/16/1993, swamp along a stream on the terrace, 07/16/1993, River basin. Koyverelyan: mouth of the river. Koyverelyan, 2nd floodplain terrace, shrubs, 07/29/1993, Tamvatney mountains: Martian mountain, gravelly-loamy trail, 07/27/1994, in the same place, spotted grass tundra with dwarf clumps at the bottom of the slope, 07/27/1994, River basin. Tamvatvaam: watershed of the Tumgytveem and Razliv rivers, sedge-mixed grass eutrophic swamp, 07/19/1994, p. Tundra at the mouth, plume of the hillside, forbs and shrubs of the tundra, 07.27.1994, Upper course of the river. Chiranay: mouth of the river. Mayolgykonvaam, nival watershed on the Pesets mountain, 07/05/1995, Chiranay mountains, raw tundra (swamps) on a plume, 07/22/1995, Sources of the river. Nygchekveem, Koryak ridge: a mixed-moss swamp on a plume, usual, 07/06/1996, Upper river. Nygchekveem: Lake Smooth, willows from S. pulchra along the hollow of the drain, 06/29/1996, Average course of the river. Nygchekveem: in the range of the mountain Nygchek and Lake. Eternal, mixed shrubs of a floodplain terrace, 07.24.1996, Outskirts of Anadyr: Mount Dionysia, raw tundra, 2.07.1993.

Carex sordida Heurck. et Mueller Agr. - Sedge dirty. Rarely, sporadically. Shrubs and meadows of floodplain terraces. Far Eastern Boreal. Our location is the most northeastern within the range of the species. Known from okr. pos. Markovo, p. Algan, river basin Penzhina and the Gulf of Corfu.Distributed from Japan and Northeast China, the Korean Peninsula and Transbaikalia to Okhotia and Kamchatka.

Middle and lower river. Great: 5 km below the mouth of the river. Chiranay, a high terrace on the left bank, coppice meadows, in alder forests, 08/17/1995, Upper course of the river. Chiranay: p. Yenattytvargyvam, shrubs of the first floodplain terrace, 07.26.1995.

Carex stans - upright sedge

Carex stans Drej. - sedge erect. Collected once. North Eurasian hypoarctoboreal bog species. Is a subspecies or even ecological race C. aquatilis Wahl. In the northern part of the Koryak Upland, it is extremely rare, only some local populations with a stretch can be attributed to this taxon. In all likelihood, the range of this species captures only the Lower Anadyr Lowland.

Oz. Mainz: west coast, coastal sandy beach in the zone of seasonal fluctuations in lake level, 07/20/1996.

Carex stilosa C.A. Mey. - sedge columnar. Seldom. Willows, meadows on floodplain pebbles, nival tundra. Beringian hypoarctoboreal species. Known from the east of the Chukchi Peninsula and Kamchatka. Dense plant with sharp narrow leaves. Scales at the base are not filamentous, light brown. The view is slightly different from C. podocarpa ssp. koraginensis, This is probably a variant of the latter associated with hyperbasites.

Tamvatney mountains: Martian mountain, nival tundra under the slope, sedge tundra, 07.26.1994, Middle course of the river. Chiranay: mouth of the river. Openwork, floodplain pebble, willows - meadows, 08/05/1995.

Carex tripartita All. var. - Three-part sedge. Everywhere in the zone of boreal dwarfs. Not rarely. Forbs of tundra, swamps, meadow nival tundra, meadows and bushes of the floodplain, river banks. Circumpolar plurizonal erosiophilic species.

In the upper reaches of the river. Chiranay, mixed shrubs of the floodplain terrace of the river. Vilyuy near Mount Gloomy there is a strange form of plants with a male underdeveloped spikelet, which is hidden among two female ones. Female spikelets, like C. tripartita, some of them at the bottom with male flowers. Forms turf. The leaves are almost equal to the stems, narrow up to 1.5 mm wide. Maybe a hybrid C. tripartita Allioni x C. cinerea Pall.

Upper river Khatyrka: ruch. Maisky, swamp along a stream, 07/16/1993, in the same place, river floodplain, 07/12/1993, River basin. Tamvatvaam: p. Tundra at the mouth, mixed grass tundra on the plume of the hillside, 07.24.1994, the middle course of the river. Tundra, icy section of the floodplain, 07/21/1994, the shore of a moraine lake at the head of the river. Spill, 07/19/1994, the middle course of the river. Tumgytvey, shrubs of the middle floodplain, 07/16/1994, Upper river. Chiranay: Ugryumaya mountain, avalanche tray, nival tundra, 07/13/1995, the mouth of the river. Elgiveem, a drainage hollow on a slope, shrubs, 07/19/1995, The middle course of the river. Chiranay: okr. Lake Good, meadow on the ledge of a high (6 m) terrace, 1.08.1995, in the same place, meadow under the ledge of a high terrace, 07/30/1995.

Carex ursina Dew. - Bear sedge. Seldom. Damp plots in the tundra along the hollows, along the banks of the Anadyr estuary, on loamy spots along the river terraces, coastal marches. North American-North Asian Arctic species. Our plants are taller (10-13 cm), spikelets 2, the final spikelet is somewhat larger. The leaves are longer than the stems and rise above the spikelets. It forms quite thick sods.

The average course of the river. Chiranay: p. A winding, moraine ridge, loamy spots on the river terrace, nival-soaked place, 2.07.1995, Outskirts of Anadyr: sea coast, 07.07.1993.

Carex vaginata Tausch. ssp. algida (Turcz. ex V. Krecz.) Galanin somb. prov. - Sedge vaginal cold. Everywhere. Infrequently. Floodplain pebbles, floodplain meadows and willows, sedge-forbs and forbs and shrubs of tundra, nival meadow tundra, alder. Circumpolar arctoalpine tundra species. Our plants are large and deviate to C. falcata Turcz. Sacks with clearly visible veins, sometimes with small papillae, or tiny short hairs.

Upper river Khatyrka: thermokarst depressions on a high terrace at the mouth of the creek. Maisky, 07/12/1993, floodplain of the Khatyrka River, 07/12/1993, intermountain hollow near Mount Evlanyr, dry mounds on the watershed, 07/20/1993, along a stream in the mountains, 07/08/1993, Middle and lower reaches of the river. Great: 5 km below the mouth of the river.Chiranay, high floodplain on the left bank, willow-meadows-coppice meadows, 08/12/1995, Tamvatney mountains: gravel-loamy plume of the Martian mountain, sedge-colored tundra, 07/27/1994, eastern macro slope of the massif, gravelly slopes, 07/31/1994, River basin . Tamvatvaam: the mouth of the river. Tundra, mountain trail, herb-shrub tundra, 07.24.1994, the middle course of the river. Tumgytveem, thickets of cedar dwarf on the side of the mountain, 07/14/1994, the mouth of the river. Tundra, a hollow of a drain on a slope, willows from S. pulchra, 07.24.1994, Upper course of the river. Chiranay: mouth of the river. Mayolgykonvaam, high (8 m) moraine terrace, 07/04/1995, the mouth of the river. Elgiveem, shrubs of a high floodplain, 07/19/1995, plume of Mount Chelka, sedge tundra, 07/19/1995, Middle course of the river. Chiranay: okr. Lake Good, low terrace, shrubby tundra, 07/31/1995, The sources of the river. Nygchekvei, Koryaksky ridge: raw meadow on a plume, 07/06/1996, mixed grass alders on a slope, 07/06/1996, Upper river. Nygchekveem: against Mount George (turn), moraine hills at the foot of the mountain 284 m, 07.24.1996, okr. Lake Smooth, moraine ridges, 2.07.1996, Surroundings of the city of Anadyr: Mount Dionisia, nival tundra on a ledge, 2.07.1993, plume of Mount Dionysia, raw tundra, 2.07.1993,

Carex vaginata Tausch. ssp. falcata (Turcz.) Galanin comb. prov. - Sedge vaginal crescent. Infrequently. Mostly alders on the slopes. Boreal East Siberian-North American forest species. In our region C. algida and C. falcata form a single morphologically continuous superpopulation.

River basin Tamvatvaam: the mouth of the river. Tundra, drain hollow, willow from S. pulchra, July 24, 1994, the average course of the river. Tumgytveem, thickets of cedar dwarf on the side of the mountain, 07/14/1994, Upper course of the river. Chiranay: mouth of the river. Elgiveem, thickets of alder and cedar dwarf on the slope of Mount Chelka, 07/20/1995, Mount Ugryumaya, alder on the slope in the lower part, 07/13/1995.

Carex vanheurckii Mueller Arg. - Sedge Van Hurk. Everywhere. Not rarely. Shrub-lichen tundra on slopes and fluvioglacial terraces, rocky talus. East Asian Montana-Boreal mountain forest species. Deviates to C. globularis. From the northern part of the Koryak Upland is known only from Lake. Mainz. There are a lot of fees from the southern part of the Koryak Upland. Our fees are the northernmost in this part of the range.

Upper river Khatyrka: floodplain, 07/12/1993, brook. Maisky, subsidence on the terminal moraine, 07/11/1993, ibid., Depressions on the moraine terrace, dominant, 07/07/1993, lichen tundra on a high terrace with thickets of elfin, 07/16/1993, River basin. Tamvatvaam: the middle course of the river. Tundra, lichen wasteland on the floodplain terrace, 07.24.1994, ibid., From S. pulchra on the hollow, 07.24.1994, Upper course of the river. Chiranay: mouth of the river. Mayolgykonvaam, high (8 m) terrace, dry canal, mixed willows 4.07.1995, Mount Ugryumaya, rocky scree on the top of the mountain, 07.13.1995, Upper course of the river. Nygchekveem: okr. Lake Smooth, moraine ridge, 2.07.1996, ibid., Shrub-motley grass tundra on a floodplain terrace, 06.28.1996, Middle course of the river. Nygchekveem: moraine under Mount Nygchek, dwarf thickets on a high hill, 08/05/1996.

Carex vesicaria - Bubble sedge

Carex vesicata Meinsh. - sedge bubbly. Everywhere. Not rarely. Swamps, muddy depressions in floodplains, gravels. East Asian-American hypoarctoboreal floodplain-bog species. From the northern part of the Koryak Upland, a collection from the upper reaches of the river was known. Khatyrka.

Upper river Khatyrka: ruch. Maisky, thermokarst depressions on a high fluvioglacial terrace, 07/12/1993, eutrophic swamp along a stream on a high terrace, 07/16/1993, intermountain hollow near Mount Evlanyr, floodplain terrace, 07/20/1993, alder on a terraced slope, 07/12/1993, Pool r . Koyverelyan: mouth of the river. Koyverelyan, silty duct in the floodplain, alder, 07/29/1993, Middle and lower reaches of the river. Great: 5 km below the mouth of the river. Chyrynay, high floodplain on the left bank, willow meadows, alder-coppices, 08/17/1993, River basin. Tamvatvaam: floodplain pebble at the confluence of the Tumgytveem and Tundrovaya rivers, 07.20.1994, Upper course of the river. Chiranay: Mount Gloomy, r. Vilyuy, swampy flood plain with channels, 07.15.1995, ibid., Swamp, 07.15.1995, Middle course of the river. Chiranay: okr. Lake Kind, a lake in the tundra, a swamp along the edge, 07/31/1995, The middle course of the river.Nygchekveem: okr. Lake Gytgykay, a dried-up lake on the moraine, 08/08/1996, in the same place, the shore of a stream flowing from lake. Gytgyky, 08/08/1996.

Carex vesicaria L. - Bubble sedge. Everywhere. Infrequently. Sedge meadows and swamps are sometimes dominant. Euro-Siberian hypoarctoboreal floodplain-bog species. Our patterns are not typical, shy to C. jacutica V. Krecz., From which they differ in light yellow-green sacs.

Upper river Khatyrka: high fluvioglacial terrace near the mouth of the stream. Maisky, thermokarst depressions, 07/12/1993, Middle and lower reaches of the river. Velikaya: 6 km above the weather station, meadow along the canal, 08/15/1995, Middle course of the river. Chiranay: okr. Lake Good, swamp along the edge of the lake, 07/31/1995, Upper course of the river. Nygchekveem: against the mountain Janranay, sedge meadow along the shore of the lake. Lowland, dominant, 07/19/1996.

Сarex williamsii Britton. - Sedge Williams. Everywhere. Infrequently. The nival tundra and meadow tundra, moist and wet tundra along the false lows of runoff. North American-Far Eastern-Siberian hypoarctoboreal montane mountain tundra species. News for the northern part of the Koryak Highlands. This is essentially a subspecies. C. capillaris L., characterized by very small sizes. Small, up to 5-6 cm in height, leaves are equal in height to generative shoots, forms sods.

Upper river Khatyrka: thermokarst depressions on the fluvioglacial terrace at the mouth of the creek. Maisky, 07/12/1993, Upper course of the river. Nygchekveem: mouth of the river. Ichguigeynei, nival valley, on the floodplain terrace above the ledge of the moraine hill, 06/30/1996, against Mount George (turn), willows from S. alaxensis along the brook on the terrace, 07.24.1996, Surroundings of the city of Anadyr: Mount Dionysia, a plume, raw tundra, 2.07.1993.

Rod Kobresia - Kobresia

Kobresia filifolia (Turcz.) C.B. Clarke = K. simpliciuscula ssp. filifolia - Narrow-leaved cobresia. Seldom. Collected once. East Siberian-West American Arctic mountain erosiophilic (?) Species. The nearest known locations are in the vicinity of the village. Ust-White.

Upper river Nygchekveem: against the mountain of George (turn), the raw hummocky tundra at the foot of the mountain 504 m, 07.24.1996.

Kobresia myosuro>

Upper river Khatyrka: intermontane depression near the Evlanyr hill, dry hillocks on the watershed, Kobresia tundra, 07/20/1993, Tamvatney mountains: low floodplain terrace of the river. Tamvatvaam, old pebble, 07/29/1994, clump of tundra with individual bushes of cedar dwarf on a high moraine terrace, 07/30/1994, hummock sedge-gunnock tundra on a slope plume, 07/27/1994, Sources of the river. Nygchekveem, Koryak Range: creek. Nakhodka, floodplain terrace, mixed grass tundra, 07/07/1996.

Kobresia sibirica (Turcz. Ex Ledeb.) Boeck. - Siberian cobresia. Infrequently. Raw tundra and mixed lawns on streams and on the saddles of the mountains. Siberian-West American arctoalpine tundra species. The nearest locations are the surroundings of the village. Ust-White. Some individuals shy away from K. simpliciuscula (Wahl.) Mackenzie.

Tamvatney mountains: raw tundra on the saddle, 07/31/1994, ibid., Mixed grass lawns along a mountain stream, 07/26/1994.

Kobresia simpliciuscula (Wahl.) Mackenzie ssp. simpliciuscula - Cobresia is simple. Sporadically in the zone of boreal dwarfs. Infrequently. Sedge-mixed grass, sedge tundra, mixed grass bushes of floodplains and floodplain terraces, rocky talus, moraine basins. Almost circumpolar arctoalpine species, gravitating to limestones. The nearest location - the surroundings of the village. Ust-White. Some individuals shy away from K. myosuroides (Viller) Fiori et Paolett - have elongated spikelets and large sizes.

Tamvatney mountains: a pass in the area of ​​Mount Shamanya, a saddle near a watershed lake, sedge-grass tundra, 07.26.1994, ibid., Loamy-gravelly plume of the Martian slope, sedge tundra, 07.26.1994, Upper river. Chiranay: the middle course of the river. Mayolgykonvaam, duct in a high floodplain, mixed willow grasses, 07/06/1995, Ugryumaya mountain, summit mixed grass tundra, 07/13/1995, mouth of the river. Mayolgykonvaam, dry shrubs of the first floodplain terrace, 07/07/1995, Chiranay mountains, clump tundra on the slopes, talus, 07/22/1995, Upper river. Nygchekveem: the vicinity of Lake. Smooth, hollow on a moraine hill, 2.07.1996.

This material uses images from electronic identifiers taken on the websites of Kamchatka Region, Chukotka Autonomous Okrug, the Polar-Alpine Botanical Garden Institute of the Russian Academy of Sciences, as well as the site "Plants of Alaska". We are grateful to the authors of the photographs.

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Abstract of a scientific article on biological sciences, author of a scientific work - F. Chetverikov

For the first time, colonies of ticks Novophytoptus rostratae Roivainen, 1947 were found in the airways of leaves, leaf sheaths and under the epidermis on the stems of plants of the family Suregasee L.These mites feed on the juice of chlorenchyma cells and cause the formation of “brown stripes”. In addition to females, eggs and nymphs of 1 and 11 ages, which were still unknown in ticks of the genus Novophytoptus Roivainen, 1947, were found inside the Osokov’s air cavities. 27 new species of host plants belonging to the genera Carex L., Eriophorum L., Bolboschoenus Palla (= Scirpus L. pro parte) and Scirpoides $> egu er (= Holoschoenus Link / Detailed descriptions and drawings of ticks of N. rostratae from the air cavities on the sedge of the inflated nose (Carex rhynchophysa S. A. Meu / are given. Some biology features are discussed wintering and migration, a hypothesized relationship between the size of the air clear cavities in the leaves of different types of sedges with the degree of occupation by their ticks.

The text of the scientific work on the theme "Some features of the morphology and biology of the four-mite no vophytoptus rostra tle Roivainen, 1947 from the Osokovs (family Cyperaceae)"

Bulletin of St. Petersburg State University. Ser. 3,2004, no. 3

SOME PECULIARITIES OF MORPHOLOGY AND BIOLOGY OF THE FOUR MILITARY MITS NO VOPHYTOPTUS ROSTRA TLE ROIVAINEN, 1947 WITH SANDLING (SEV. CYPERACEAE)

Introduction Currently, in the suborder Tetrapodili Bremi, 1872 (superfamous Eriophyoideae), 5 families are distinguished: Pentasetacidae Schevchenko, 1991, Nalepellidae Roivainen, 1953, Phytoptidae Murray, 1877, Eriophyidae Nalepa, 1898, and Diptilomiopidae. developed forms, combining more than 95% of the described species of four-legged ticks. This study continues a series of works 2,3,6,7 devoted to the study of the most ancient representatives of Tetrapodili, which are traditionally considered species of the families Pentasetacidae, Nalepellidae and Phytoptidae. Compared with the giant family Eriophyidae (2918 species), the volume of these archaic groups is small: they include 1.68 and 54 species, respectively. Flares of the families Pentasetacidae and Nalepellidae were found only on gymnosperms from the Pinopsida class. The reasons why the ancient nalepellids and pentasetacids were preserved on conifers are more or less clear and are associated primarily with antiquity and conservatism of the signs of their owners.

Among the ancient families of four-legged ticks, a special place is occupied by this. Phytoptidae, which currently includes 8 genera and 42 species. Most of its representatives (25 species) are associated with monocotyledonous plants of the orders Arecales, Liliales, Juncales, Poales and Cyperales. The other 17 species are inhabitants of the dicotyledonous subclasses Hamamelididae, Rosidae, and Dilleniidae. Within the Phytoptidae family, two life forms of ticks are found. One of them is characteristic of species living on the open surface of plants, mainly palm trees, and is represented by ticks with a compact, small-ringed body, a large, often smooth shield and a gnatosome directed downward. Ticks of the genera Acatrix K., Mackiella K., Propilus K., Protrix K., and Retracrus K. belong to this group. Another life form is characteristic of ticks living in enclosed spaces and is represented by species with a many-ringed body, an elongated gnatosome and a small one a shield on which there is a drawing. Mites of the genera Phytoptus Murrey, Anehyphytoptus K. and Novophytoptus Roiv have such a morphotype. Along with the adaptive characters that characterize the morphotypes of hidden and open-living ticks, fi-toptids, like other ancient Tetrapodili, are characterized by plesiomorphic characters. These are considered to be the presence of s.f. and / or sdl on the dorsal scutellum, the presence of sub-dorsal setae and spurs on the tibia. “Among phytoptids, the species P. pytoptus and Osokovy, grouped in the cariéis group, have the most complete set of ancestral characters. These mites are believed to be along with representatives of p Trisetacus Keifer (family Nalepellidae) with conifers is the closest to the original forms for Tetrapodili, the reasons why the ancient figoptids were preserved precisely on sedges are not yet known.

This work is devoted to the study of ticks p. Novophytoptus with sedge. Species of this genus are very similar in appearance to representatives of p. Phytoptus from the cariéis group. However, compared with them in ticks p. Novophytoptus observed loss of s.sd., s.fem. and spurs on the legs of the first pair, legs, as well as a significant (8–9-fold) increase in the length of s.d.2 and lengthening of the body in the middle part, which caused the epiginium to be moved away from the coxosternal skeleton. Based on these signs, X. Roivainen (N. Roivainen) in 1953 identified as part of this. Phytoptidae tribe Novophytoptini Roiv. Currently, it includes six species: Novophytoptus rostratae Roiv. and N. tauricus Mitrophanov, Sharonov, Sekerskaya with sedges, N. stipae K., N. glyceric¡е Scoracka et Boczek and N. ammophyllae Scoracka et Boczek from cereals and N. sitvai Flechtmann with salmon.

Literature data on ticks p. Novophytoptus is extremely scarce. Descriptions of famous to

About F.E. Chetverikov, 2004

At present, species are given for a few females and single males.There is no information on the findings of nymphs I and II and eggs in the literature. One find of these ticks is known in North America, one in South America and five in Europe (Table 1). Five out of six species of the river. Novophytoptus were collected from the leaf surface, and only one (N. silvae Flecht- ,, mann, 2004) was also found in spikelets and in the spaces under the epidermis of old leaves. H No visible damage caused by ticks of the p. Novophytoptus was not detected.

Table 1. Finds of pincers p. Novophytoptus Roivainen., 1947 (according to the literature). city ​​„R

Type of tick Literary source Place of find Date of find Host plant Localization, 1 on rzhgteniya

, N. ammophilae Skoracka, Boczek Poland 07.21.1999 Ammophila arenaria (L.) Link. on the surface of the leaves

N. glyceriae Skoracka, Boc-zek Poland 07/27/1999 Glyceria maxima. (Hartman) Holmb. on the surface of the leaves

N. rostratae Roivainen Finland 08.16.1944 Carex rostrata Stok. (C. inßata Sut, C. ampullacea Good.j in the median leaf groove

N. rostratae? Roivainen Spain 07/11/1953 Carex flacca Schreb. (C. glauca Scop.,) Inside the leaf sheaths

08/09/1953 Scirpus sp. in leaf grooves

N. silvai Flechtmann Brazil 10/07/2003 Cyperus gyganticus Vahl. in leaf grooves, spikelets and under the epidermis of old leaves

N. stipae Keifer USA, California. 04/11/1962 Stipa speciosa Trin. et Rupr. inside twisted leaves

N. tauricus Mitrofanov, Sharon, Secerskaya Ukraine, Crimea 01/20/1977 Carex sp. not indicated

.. A type species of the genus N. rostratae Roivainen was first found in 1944 in a leaf groove on a sedge swollen (Carex rostrata) in Finland and was described in 1947 by Roivainen. To date, this species was considered one of the rarest among the phytoptids that live on sedges. Royvainen reports that for several years (1944-1953) he unsuccessfully tried to re-find ticks of N. rostratae. Only once, nine years after the first find, he managed to find this species in Spain on the plants Carex flacca and Scirpus sp. in the median leaf groove. He also indicates that he did not have the opportunity to describe in detail and accurately determine the ticks found due to insufficient material, so in the publication after the species name he put a question mark: “N. rostratae? ” l

Since 1985, we have been collecting ticks from the Osokovs. Moreover, representatives of the river. Novophytoptus were found singly on the upper surface of sedge leaves. In July 2002, while collecting material in the Plesetsk district of the Arkhangelsk region, looking under the binoculars for the stems and leaves of the sedge C: juncella, we noticed moving ticks under the brown epidermis of the leaf sheath. After removal of the epidermal layer in the spaces between the vascular bundles, female tick mites were found. Novophytoptus, their eggs, nymphs of I and II age.

In the future, the search for ticks continued not only on the surface of the leaves, but also under the epidermis. The result was not slow, affect. Soon new mite colonies were found inside C. juncella leaf blades. It turned out that in places of congestion

ticks of the parenchyma and epidermis cells were brown. From the surface, these patches looked like brown stripes. This observation significantly facilitated the collection of ticks, since the damage caused by them (brown stripes) is visible to the naked eye. In the end, colonies of ticks p. Itor and Orshv were found on more than twenty forks of plants of this family. Suregasee.

Material. The material for the study was the collection of ticks of the river. Ys ^ orkuYurShB from the North-West of Russia (Arkhangelsk region, Plesetsk district, Leningrad region from the Suyai river to the northern Ladoga), produced in 1985, 1995 and 2001-2003. In 2002-2003 ticks were also collected in the south of Ukraine (from Dnepropetrovsk to the northern coast of the Sea of ​​Azov). During this time, we examined 43 species of plants of this family. Suregaseae, and only 27 of them were found ticks of the river. Msmorku JurSh (tab. 2).

Table 2. Host plants of ticks p. IochorIuYuRSh Noshashep, 1947 in the North-West of Russia and the south of Ukraine

North-West Russia South Ukraine

Arkhangelsk region Leningrad region. Dnepropetrovsk, Zaporizhzhya region

1. Carex acuta L. (= C. gracilis 1. Carex acuta L. (

C. gracilis 1. Bolboschoenus maritimus (L.)

Curt.) Curt.) Palla

2. C. cespitosa L. 2. C. cespitosa L. 2. Carex acuta L (-C. Gracilis

3. ' C. rostrata Stok. (- C. inflata 3. C. rostrata Stok. (- C. Inflata CilTt)

Sut, C. ampullacea Good) SuL, C. ampullacea Good) 3. C. atherodes Spreng

4. C. rhynchophysa C. A. Mey 4. C. rhynchophysa C. A. Mey 4. .C. colchica s.sp. colchica Egor.

5. C. limosa L. 5. C. limosa L. 5. C. contigua Hoppe

6. C. vesicaria L. 6. C. vesicaria L. 6. C. distans L.

7. C. elata All. . 7. C. disticha Muds 7. C. externa Good

8. C. flava L. 8. C. pallescens L. 8 C. hirta L.

9. C. juncella (Fries) Th. Fries 9. C. panicea L. 9. C. praecox Jacq.

(= C. juncea Fries) 10. Eriophorum vaginatum L. 10. C. secalina Willd. ex Wahlenb.

10. C. lasiocarpa Ehrh. 11. C. stenophylla Wahlenb.

Ik C. nigra (L.) Reich 12. Scirpoides holoschoenus (L.)

(= C. goodenowii Gay) Sojak.

12. Eriophorum vaginatum L.

13. E. polystachyum L.

The main material was collected from under the epidermis of the leaves, leaf sheaths and stems. Single ticks were found on the upper surface of leaf blades and spikelets.Plants were examined under the binocular, the four-legged ticks found were transferred to the Fora-Berlese liquid, and microscopic preparations were made. The repara ™ was viewed under an MB-3 microscope with a KF-4 phase-contrast device and an * 90 immersion lens.

All ticks are assigned to p. Novophytoptus. Measured 6 samples of 12 ticks from different plants from this. Suregasee (Bolboschoenus maritimus, Carex limosa, C. rhynchophysa (2 samples), Eriophorum pofystachyum and E. vaginatum). For a number of features, a pairwise comparison of the samples by the t-test was carried out.

Unpublished data on the anatomy and biology of sedges of the Leningrad Region from student thesis by T. B. Sokolovskaya (1955) were used (with permission of the leadership of the Department of Botany of St. Petersburg State University).

Results and discussion. Of the 27 plant species on which p. Novophytoptus, only one (Carex rostrata) was previously known as the host of Novophytoptus rostratae. On 22 species of sedges and plants from the genera Bolboschoenus Palla (= Scirpus L. pro parte), Eriophorum L, and Scir-po>

For the north of Europe, only one species p is known so far. Novophytoptus - N. rostratae. The description of this species, as well as the figures given in it, are not detailed enough, therefore, the species N. rostratae

in rewriting. Unfortunately, the type material has not been preserved, and it is not yet possible to obtain and describe a neotype. The most representative material for this species was obtained from sedge of a bloated nasal (C. rhynchophysa). This morphology, biology and systematic position of this sedge is close to the sedge swollen (C. rostrata), with which the species N. rostraíae was described. Since the literature does not contain data on the morphology of the species N. rostraíae (except for the description of the species), we give below a description of the mites found under the epidermis of the sedge swollen.

Novophytoptus rostraíae Roivainen, 1947. Worm-shaped ticks, dull white. The body length of the female is 241.5 (220-275) ', the height is 54.0 (50-60). The length of the flap is 38 (36.5-40). The shield pattern (Fig. 1, C) is represented by five longitudinal lines: a clear medial line and two admedial lines and a pair of thin submedial lines. The medial line is incomplete, expressed only in the posterior half of the scutellum. Admedial lines are thicker than medial. At first they diverge slightly, and then gradually approach and, almost merging, reach the front edge of the flap. As a result, a figure resembling a three-toothed fork is formed. The submedial lines are thin, barely noticeable, starting from the tubercles of s.d.2 and, curving in an arcuate manner, merge with the medial line in the anterior third of the scutellum (see Fig. 1, B). Lengths. d. 1 -13.5 (10-16), s. d. 2-105.5 (94-118). Gnagosome 25.0 (24-26.5) directed forward.

Fig. 1. Novophytoptus goIga1ae Ihmuatep, 1947 female. A - side view, B - coxo-sternal skeleton and epiginium, C - drawing of the scutellum, D - empody: / ​​- legs I, 2 - legs II, D - limbs: 1 - leg 1,2-leg II.

All measurements are given in micrometers.

Leg length I - 37.0 (31-40), tibia - 10.5 (8-11), s.tib - 4.0 (4-5), tarsus - 5.5 (5-6). Claw - 5.0 (4.5-5) with a small washer. 4-ray empody, two distal rays more than three times longer than proximal. Leg length P - 36.0 (31-38.5), tibia - 8.5 (7.5-10), tarsus - 5.0 (5-6). Claw - 8.5 (8-10) with a small washer. The empathy of the legs P is 4-beam, asymmetric: one of the two distal rays is more than three times longer than all the others (Fig. 1, D, D). Chest ina T-shaped. Cokes are streaked with longitudinal, wavy lines. The length of the epiginia is 6.5 (6-6.5). The cap of the epiginia is smooth (Fig. 1, S). Length s. gen. -15.0 (12.5-18).

The opistosome is equally ringed, with 78 (73-83) dorsal half rings. Both the dorsal and ventral half rings are covered with oval micro-tubercles. Between the coxo-fern skeleton and epiginium 11 (10-12) ventral semirings. The caudal part of the body is saccularly expanded (Fig. 1, A). The last 6-7 rings of opistosomes are dorsally smooth, covered by micro-tubercles elongated along the body axis. Length s. 1-23.0 (20-30), s. v. 1-26.5 (20-33.5), s. v. 2-13.5 (11.5-16), s. v. 3-18.0 (16-20), s. acc. - 6.0 (5-6.5). Setae s.v.3 distinct, acicular pointed, 1.5–2 times thicker than other setae of the opistosome. The number of half rings to s. 1 - 9 (8-10), the number of half rings between s. 1 and s. v. 1-14 (12-17), between s. v. 1 and s. v. 2-17 (16-18), between s. v. 2 and s. v. 3-30 (27-33).

Telosome includes 7 rings.

Measurements are given in 12 copies.

The host plant is Carex rhynchophysa S. A. Meu.

Localization on the plant - inside the leaves (in the air cavities), mite-populated areas of the leaf are visible as brown stripes.

Date and place of collection - 07/15/2003, Leningrad Region, Pavlovsk, park.

Males: not found.

The colony was attended by numerous nymphs I, II and eggs. •

During the summer, we met mostly unpainted, matte white females. However, in spring and early autumn, almost all the females in the samples were orange-red.Thus, on 10.092002 immovable, orange-red females, bound in tangles of 10-15 individuals, were found in the park of Pavlovsk in the Leningrad Region (in the same place where the unpainted females were described) under the epidermis of the Sages rhynchophysa leaf. The same colored females laying eggs were found in spring (05/18/2003) in the village. Mozhaysky Gatchinsky district of the Leningrad region under the epidermis of the sedge biaxial leaf (C. disticha). We suggested that colored ticks are the winter form of N. rostratae females, and unpainted ones are summer. To find out if there are morphological differences between them, a sample of wintering orange ticks found in the fall in Pavlovsk was measured. The obtained data were compared by the ¿criterion with measurements of the white females described above (Tables 3, 4). It turned out that winter and summer females significantly differ in the length of the gnatosome, epiginia, legs I, and setae s.d. 1, s.d.2, s.v. 1, s.v.2 and the number of dorsal semirings.

Table 3. Characteristics by which ticks of N. goyashgay Yoguatep, 1947 from different populations with pairwise comparison of samples (1,2,3,4,5 *) significantly differ by the ¿-criterion (p Can't find what you need? Try the literature selection service.

Original Roivainen data, -1947

Carex rhynchophysa S.A. Meu C. limosa L. Eriophorum polystaehyum L. E. vaginatum L. Carex rostrata Stok.

1 1 2 s L_ 4 i 5 Description

Sign Date of discovery

15.07.2003 | 10.09.2002 | 22.07.2002 • 22.07.2002 1 22.07.2002 - | 16.08.1944

Pavlovsk, Leningrad region Arkhangelsk region, Plesetsk district Finland

Localization on the plant

inside the air cavities on the leaves inside the air cavities • on the leaves in the middle groove of the leaves inside the air cavities on the leaves and stems inside the air cavities on the leaves and stems in the middle groove of the leaves

Color matte white orange red matte white matte white matte white white

Body length 241.5 (220-275) 257.5 (206-294) .234 (175-268) 232.2 (206-265) 238 (220-268). 200-240

Gnatosome length, ¡25.0 (24-26.5) 27.5 (26.0-28.5) 25 (23.5-26.5) 26.0 (24-28.5) 25.0 ( 22-28) 28

Length s.d. 1 13.5 (10-16) 16 (11.5-18.5) 13.0 (12.5-13.5) 13.0 (12.5-13.5) 14.0 (12.5 -16) 12

Length s.d.2, 105.5 (94-118) 114.0 (100-125) 81.0 (56-105) 112.5 (100-130) 119.5 (100-130) 90

The number of dorsal 1 78 (73-83). 74 (68-77) 73.5 (67-78) 73 (68-77) 70 (67-78), "72

Leg length I 37.0 (31-40) 41 (38-43.5) 36 (35-37.5) 39.0 (37.5-41) .38.5 (37.5-40) 38

Tibia I length 10.5 (8-11) •• 11 (10-12) 9.5 (8.5-10) 10.5 (10-11.5) 11.0 (10-12) 11

The length of the epiginia is 6.5 (6-6.5). 7.5 (6.5-8) 6.5 (6.5-7.5) 6.5 (6-8.5) '6.5 (6-7) 9.:

Length s.v. 1 26.5 (20-33.5) 31.0 (26.5-37) 29.5 (20-40) 22.5 (17.5-26.5) 24.5 (22.5-26 ) 30-35 ..

Length s.v.2 13.5 (11.5-16) 16.5 13.0 (10-14) 13.5 (12-17.5) 15.0 (12.5-18.5) 14

Length s.v.3 18 (16-20) 19 (15-23.5) 18.0 (15-20) 18.0 (16-20) 19.5 (17.5-23) 23-25

Length s.acc. 6.0 (5-6.5) 6.5 (5.5-7.5) 5.5 (5.0-6.5) 5.0 (5-6) 5.5 (5-6) • 5

When comparing our data with the description of the species N. rostratae, it turned out that the mites we collected were characterized by longer sdl and s.d2 setae, had a greater number of dorsal half rings, shorter opistosome setae, shorter leg length I, and epiginium cap (see table. 4). In the description of the species N. rostratae, there are no data on the structure of empodia. In our case, the empody on legs I and II differed in shape (see Fig. 1> D, D). Royvainen does not mention the presence of submedial lines on the shield, nor are they shown in his drawing. These lines are very weakly expressed, often they were very clearly visible even at very high magnification (15x90), and in some individuals they were not visible at all.

In addition to ticks from sedge of an auriferous sediment from the Leningrad Region, we measured 3 samples, also assigned to the species N. rostratae. These are ticks found under the epidermis of leaves on the sedge of marsh (Carex limosa L.) and cotton grass (JEriophorwn potystachyum L, E. vaginatum L) in the Arkhangelsk region. Signs by which there were any noticeable differences are given in table. 4. When pairwise comparing all the samples with each other, these signs revealed significant differences (see Table 3). Most often, the samples differed in the lengths of the scutes and opistosomes. Winter and summer females with C. rhynchophysa, as well as summer females from different species of cotton grass, differ in 8 ways. All other samples have fewer significant differences (from 4 to 7). Thus, the intrapopulation seasonal variation in ticks of N. rostratae is comparable with the inter-population one.

Royvainen believed that p. Novophytoptus, living on the Osokovyh, lead an open lifestyle and are extremely rare 10.11. The discovery of a large accumulation of ticks of N. sitvai in the spaces under the epidermis inside the old leaves of the saty (Cyperus giganticus Vahl) at the end of 2003 in Brazil casts doubt on the truth of Roivainen's assumption. In the course of our study, it was found that ticks of the river.Novophytoptus from Osokovyi is widely distributed BecbMá and is usually found under the epidermis inside the leaves, stems and leaf sheaths of its host plants.

The anatomical structure of the leaves of plants of the family. Suregaseae is characterized by the presence of air cavities lined with parenchyma cells 1,4 inside. These cavities are located under the epidermis between adjacent vascular bundles along the entire length of the sheet, forming peculiar tunnels, blocked in some places by transverse strands of mechanical tissue. As a result, chains of closed chambers filled with air are formed inside the leaves of sedges. It is in these chambers that ticks of the river penetrate. Novophytoptus, where they feed on the juice of chlorenchyma cells, actively multiply and form large colonies. As a result of the activity of ticks, sections of necrotic parenchyma appear between the vascular bundles. In most cases, they are visible to the naked eye as black-brown, brown or rust-colored strips of 3-15 mm in length and 1-2 mm in width, located between adjacent veins (Fig. 3, A). Outwardly, these strips are very reminiscent of damage caused by rust fungi. In this case, we are faced with a new type of damage not known so far.

Fig. 2. Damage caused by ticks of the genus Novophytoptus Roivainen, 1947 (scheme, scale not observed).

A is the appearance of the area of ​​the leaf populated by ticks: I is the “brown stripes”, B is the colony of ticks of N. rostratae inside the leaf: / is the epidermis, 2 is the parenchyma, 3 is the airway.

. Fig. 3. Fragments of leaf blades of sedges with air cavities (/) (unpublished data by T, B. Sokolovskaya, 1955).

A - Carexpallescens L., B- ^ C. acuta L., B —C. rhynchophysa S.A. Meu.

Tick ​​infection p. Novophytoptus of different species of host plants varied greatly. Thus, on the sedges of C. extensa, C. pallescens, C. panicea, and C. stenophylla, single mites of p. Novophytoptus only on the surface of intact leaf blades. In most cases (at Sageh atherod.es, C. cespitosa, C. colchica, C. contigua, C. distans, Carex disticha, C. hirta, C. lasiocarpa, C. limosa, C. nigra, C. rhynchophysa, C . rostrata and C. vesicaria) we found ticks inside the air cavities on the leaves, and in the upper third of the leaf blades they were more common than in the middle and lower. In some cases, plants were infected with ticks especially severely. So, on Sageh juncella, C. rhynchophysa and Bolboschoenus maritimus, mites were found in a multitude inside the air cavities of leaves and leaf sheaths, on Eriophorum polystachyum - inside the air cavities of leaves and under the epidermis of stems, and on Sageh acuta - in all of the listed habitats.

In our opinion, the difference in the degree of infection of different species of sedges is due to the fact that the airways, where ticks multiply p. Novophytoptus, unevenly expressed in different representatives of this family. Suregasee. So sedges, on which we did not find any damage and found only single ticks, for example C. pallescens, had rather rigid, narrow leaves with weakly expressed cavities inside (see Fig. 3, A). In most cases, ticks have been found on plants with relatively thick, juicy leaves and large airways. The largest number of air cavities inhabited by large colonies of ticks was found on the sedge acute (C. acuta) and sedge swollen (C. rhynchophysa). These species of sedges grow in humid habitats and have giant air cavities (see Fig. 3, B, C).

During the collection and processing of the material, some data has accumulated regarding the status of tick colonies p. Novophytoptus. On average, 30-40 females were in one air cavity. Eggs, nymphs I and II were found only inside the air cavities; neither eggs nor nymphs were open on leaf blades. The largest number of eggs (up to 20 in one cavity) and nymphs of both ages (up to 25 in one cavity) were found on sedges in the Arkhangelsk region in the second half of July. In Ukraine, by the beginning of August, populations of N. rostratae were mainly represented by adult females and a few nymphs II. In Leningrad Oblast, by September, populations consisted of only adult females and single nymphs II. Males were not found in any sample.

In the second half of July in the Arkhangelsk region, during the collection of material, we observed how dozens of N. rostratae ticks crawled out of the airway through a crack in the epidermis of the C. limosa sheet for dozens and moved in different directions along the upper surface of the leaf blade. A similar fact was noted by K. Flechtmann on the example of the species N. silvai. He saw how these mites in mass crawled out from under the epidermis of old leaves of a giant giant (Cyperus giganticus) through a round hole. The penetration of the tick of N. rostratae under the epidermis was noted by us twice: 07/07/2002 on sedge of marsh ^ (C. limosa) and 07/24/2003 on sedge of swollen (C. rostrata) in the Arkhangelsk region. In both cases, single ticks penetrated into the sheet through cracks in the mechanically damaged epidermis. Flechtmann suggests that stomata can be used for tick penetration into subepi-dermal spaces.

Several times we managed to find wintering ticks of the river. Novophytoptus (Table 5). In all cases, the mites were bound into tangles of 10-15 individuals, motionless and localized in the peripheral parts of the airways of the leaves and leaf sheaths of the sedge. They were all orange-red. Finding wintering colored ticks p. Novophytoptus inside the leaves suggests that they use the same air cavities for wintering in which they live during the summer. Probably, in late spring, ticks leave them and populate new cavities on young leaves or stems. During the summer, active egg laying and the development of nymphs 1 and II occur inside the airways. Possibly, ticks also have a resettlement migration, which occurs in the middle of summer, when we observed ticks in the mass leaving the air cavity.

Table 5. Finds of wintering ticks p. Novophytoptus Roivainen, 1947 on sedges in the Leningrad Region

Date of find Location of the host plant Localization on the plant

10.09.2002.5.10.2003 Pavlovsk park Carex rhynchophysa S. A. Meu inside the airways of dry leaves

09/28/2003 ■ pos. Mozhaysky, the shore of the lake. Duderhof S. disticha Huds. in the air cavities of leaf sheaths on a drying plant

10/10/2002 Art. Marienburg, Menagerie Park, Miracle Polyana S. panicea L. in the airways of leaf sheaths on a green plant

Conclusions. Genus A ovophytoptus Yauhchatgp, 1947 from the Osokovs proved to be common in the examined territories of the North-West of Russia and Ukraine. Taking into account the fact that most of the year ticks of this genus are located inside the air cavities, it becomes clear why they could not be found for a long time and were considered one of the rarest open-living phytophthids that live on the Osokovye. With the discovery in the summer of 2002 of ticks N. gosl1ae inside the organs of host plants, it became clear that they are specialized inhabitants of the airways of leaves, stems and leaf sheaths of plants of the family. Suregasee. The brown stripes on the leaves, stems and in the leaf sheaths of the Osokovs, the formation of which are provoked by ticks of N. goiuye, are very similar to the damage caused by rust fungi. Perhaps, it is precisely for this reason that they have not yet been discovered. Maybe other representatives of the river. Novophytoptus, known to date, lead a hidden lifestyle and live inside natural spaces in the tissues of host plants. Further research is needed to clarify this issue and refine biology data.

The author sincerely thanks the staff of the Research Institute of St. Petersburg State University I. G. Bagnyuk, S. I. Sukharev and V. G. Shevchenko for discussing the earlier versions of the article and constructive criticism, and the associate professors of the Department of Invertebrate Zoology of St. Petersburg State University A. A. Dobrovolsky and A. N. Ostrovsky for discussion articles in the process of its preparation, as well as employees of BIN them. V.L. Komarova G. Yu. Konechnaya and T.V. Egorova, 6th year student of the Department of Botany of St. Petersburg State University P.G. Efimov, Entomologist of the Central Center for Social Protection of the Plesetsk District, Arkhangelsk Region. T. G. Chetverikova for help in collecting the material and identifying the Osokovs, especially thanks the staff of the Department of Botany of St. Petersburg State University for the opportunity to work in the Herbarium and use the unique library of the department.

Article recommended by prof. A. I.Granovich.

Tchetverikov F. E. Some features of biology and morphology of eriophyid mite Novophytoptus rostratae Roi-vainen, 1947.

Colonies of mites Novophytoptus rostratae Roivainen, 1947 were found within air-cavities of the leaves and leaf-sheaves and under ep>

1. Alekseev Yu. E. Osoki. M., 1996. 2. Bagnyuk I. G., Sukhareva S. I., Shevchenko V. G. Analysis of the genus as a key one in the family Trisetacus Keifer, in the family Nalepellidae Roiv. (Acari, Tetrapodili) // Acariña. 1995. Vol. 3, No. 1-2. P. 31-65. 3. Bagnyuk I. G., Sukhareva S. I., Shevchenko V. G. The main directions of evolution of four-legged ticks as a specialized group of organisms using the example of the families Pentasetacidae Shev., Nalepellidae Roiv. and Phytoptidae Murrey (Acari, Tetrapodili) // Acariña. 1998. Vol. 6, No. 1-2. P. 59-76. 4. Egorova T. V. Osoki (Carex L.) of Russia and neighboring states (within the former USSR). SPb., 1999. 5. Mitrofanov V.I., Sharonov A.A., Sekerskaya N.P. Toward cognition of four-legged ticks (Acari, Eriophyoidea) from sedge of Crimea // Zool. Vestn. 1983. No. 4. P. 81-82. 6. Sukhareva S. I. Family Phytoptidae Murrey, 1877 (Acari, Tetrapodili), its composition, structure and possible evolutionary paths // Acariña. 1994. Vol. 2, No. 1-2. P. 47-71. 7. Shevchenko V. G., Bagnyuk I. G., Sukhareva S. I. A new family of four-legged mites Pentasetacidae (Acariformes, Tetrapodili) and its significance for the interpretation of the origin of the group // Zool. journal 1991.V. 70. Issue. 5. S. 47-53. 8. Atppef W. Keys to the world genera of the Eriophyoidea (Acari, Prostigmata). USA 1996. 9. Flechtmann, C. H. W. Eriophyid mites (Acari: Eriophyoidea) from brazilian sedges (Cyperaceae) II Internet. J. Acarology. 2004. Vol. 30, No. 2. 10. Keifer H. H. Eriophyid studies B-8. Bul. Entomol. Cal. Dept. Agr. 1962. P. 1-20. 11. Roivainen H. Eriophyid News from Finland // Acta Entomol. Fennica. 1947. Vol. 3. P. 16. 12. Roivainen H. Some gall mites (Eriophyoidea) from Spain // Archivos del Instituto de Aclimatación. Almería. 1953. Vol. 1. P. 1--43. 13. Skoracka A., BoczekJ. Novophytoptus giyceriae sp. n and N. ammophilae sp. P., the new species of eriophy-ioid mites (Acari-: Eriophyoidea) from grasses // Acariña. 2000. Vol. 8, No. 2. P. 95-101. 14. Smith I. M. A new species of Eriophyid mite with eye-like structures, and remarks On the Genus Phytoptus (Acari: Prostigmata: Phytoptidae) // Canad. Entomol. 1977. Vol. 109. P. 1097-1102.

. The article was received on March 18, 2004.

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